Family Tetragonicipitidae

Kunz (1984c) reviews the family and discusses its phylogeny. He places Fearia as a synonym of Tetragoniceps and declares Phyllopodopsyllus pirgos to be a juvenile male of P. thiebaudi.

Kunz (1984a, b) describes five new species of Phyllopodopsyllus (angolensis, petkovskii, geddesi, gertrudi, mielkei, with a subspecies m. californicus) and two new subspecies of P. longipalpatus — l. madagascarensis and l. hawaiiensis.

Pallares (1982a) redescribes Phyllopodopsyllus mossmani and places P. paramossmani in its synonymy.

Oniscopsis inabai Kitazima, 1983 is added to the genus.

These additions and amendments, together with the fact that the present key does not adequately deal with sexual dimorphism in Phyllopodopsyllus, have led me to construct the following new keys to the family to replace KGG 1 (pp. 131-133).

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KGG 1 - characters

1.	Cephalothorax ornamentation p = process at posterior lateral corner present a = process at posterior lateral corner absent
2.	A.1, large dentiform projection (Note: usually similar in both sexes, but may be reduced in the male and well developed in the female) a = absent 1 = present on first segment 2 = present on second segment
3.	P.1 Enp. n = number of segments
4.	P.2-P.4., distal (or only) segment n:n:n = number of setae and spines on P.2, P.3 & P.4
5.	P.2-P.4 Exp.3 n:n:n = number of setae and spines on P.2, P.3 & P.4

KGG 1

Cph. orn.	A.1 proj.	P.1 EnP.	P.2-P.4 EnP. distal seg. setae	P.2-P.4 Exp.3 setae
p	1	2	3:3:2	6:6:5	Laophontella typica
p	1	2	3:3:1	6:6:5	L. armata
p	1	2	3:3:1	6:6:4-5	L. horrida
a	1	2	4:4:4	6:7:7	Tetragoniceps prima
a	1	2	3:3:3	6:5:7	T. bergensis
a	1	2	3:3:3	5:5:5	T. bookhouti ♀
a	1	2	3:3:3	5:4:7	T. truncata
a	1	2	3:3:3	4:4:5	T. malleolatus
a	1	2	3:3:?	6:5:4	T. longicaudata
a	1	2	?:3:3	?:5:6	T. scotti
a	1	2	2:3:3	5:5:5	T. bookhouti ♂
a	1	2	2:2:3	4:3:5	T. brownei ♀
a	1	2	2:?:3	5:?:6	T. arenicolous
a	1	2	?:?:2	?:?:7	T. dubius
a	1	2	?	?	T. brevicauda
a	2	3	3:3:2	5:4:5	Protogoniceps
a	2	2	2-3:1-3:1-3	4-5:4-6:4-7	KGG 100
a	2	2	?	?	Phyllopodopsyllus minor
a	a	3	2:2:2	4:4:4	Pteropsyllus
a	a	3	3:3:4	4:4:7	Diagoniceps trifidus
a	a	3	3:3:3	5:5:8	D. menaiensis
a	a	2	4:4:4-5	6:7:8	D. bocki
a	a	2	4:4:4	6:7:8	D. kunzi page 10
a	a	2	3:3:3	5:6:8	D. monodi
a	a	2	2-3:2-3:2-4	4-5:4-6:5-7	KGG 200
a	a	2	2-3:1:1	3:2:2	Oniscopsis
a	a	2	2:2:3	4:3:5	Tetragoniceps brownei ♂
a	a	2	?	?	Phyllo podo psyllus tristanensis

KGG 100-200 - characters

(The same five characters are used in both KGG)

1.	P.2 Enp.2 n = number of setae (Note: In the male the "setae" may include a long straight apophysis)
2.	P.3 Enp.2 n:n = number of setae in ♀ and ♂
3.	P.4 Enp.2 n:n = number of setae in ♀ and ♂
4.	P.2-P.3 Exp.3 n:n = number of setae
5.	P.4 Exp.3 n:n = number of setae in ♀ and ♂

KGG 100

P.2 Enp.2	P.3 Enp.2 ♀:♂	P.4 Enp.2 ♀:♂	P.2-P.3 Exp.3	P.4 Exp.3 ♀:♂
3	3:3	3:3	5:6	6:6	Phyllopodopsyllus chavei
3	3:3	3:3	4:4	5:5	P. medius
3	3:3	3:2	5:6	7:7	P. bermudae
3	3:3	3:2	5:6	7:6	P. setouchensis
					P. mielkei s.str.
3	3:3	3:2	4:4	7:7	P. danielae
3	3:3	3:2	4:4	7:6	P. simplex
3	3:3	3:2	4:4	6:6	P. pauli
					P. opisthoceratus
3	3:3	2:1	5:6	6:6	P. minutus
3	3:2	3:2	5:6	7:7	P. parafurciger
					P. curtus
3	3:2	3:2	5:6	5-7:6	P. furciger
3	3:2	3:2	5:6	6:6	P. longicaudatus
3	3:2	3:2	4:4	6:6	P. borutzkyi
3	3:?	3:?	5:6	6:?	P. mielkei californicus
3	3:?	3:?	4:4	6:?	P. laticauda
3	3:?	2:?	4:4	6:?	P. bahamensis
3	2:3	3:2	5:5	5:5	P. langi
3	2:2	2:2	5:6	7:7	P. parabradyi page 11
3(2♂)	3:2	3:2	5:6(5♂)	6:6	P. bradyi
2	3:3	3:2	4:4	6:6	P. paraborutzkyi
2	2:2	3:2	4:4	4:4	P. geddesi
2	2:2	2:2	4:4	5:5	P. hermani

KGG 200

P.2 Enp.2	P.3 Enp.2 ♀:♂	P.4 Enp.2 ♀:♂	P.2-P.3 Exp.3	P.4 Exp.3 ♀:♂
3	3:3	3:4	5:5	7:7	Diagoniceps laevis
3	3:3	3:3	4:4	7:6	Phyllopodopsyllus berrieri
3	3:3	3:2	5:6	7:6	P. briani
					P. petkovskii
3	3:3	3:2	5:6(5♂)	7:6	P. angolensis
3	3:3	3:2	5:5	7:6	P. thiebaudi
3	3:3	3:2	5:5	7:5	P. gertrudi
3	3:3	3:2	4:4	7:6	P. punctatus
3	3:3	3:2	4:4	6-7:6	P. mossmani
3	3:?	3:?	5:6	6:?	P. aegypticus
3	3:?	3:?	4:4	7:?	P. hibernicus
					P. laspalmensis
3(2♂)	3:3	3:2	4:4	6:6	P. hardingi
2	2:3	2:2	4:4	6:6	P. biarticulatus
2	2:?	3:?	4:4	6:?	P. xenus
2	2:?	2:?	4:4	6:?	P. paraxenus
					P. longipalpatus sstr.
2(3♂)	2:2	2:2	4:4	7:5	P. l. madagascarensis
2(3♂)	2:2	2:2	4:4	7:6	P. l. hawaiiensis