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Manual of the New Zealand Flora.

4. Eleocharis, R. Br

4. Eleocharis, R. Br.

Stems simple, erect, without perfect leaves. Sheaths few. the uppermost cylindric, truncate or produced on one side into a short tooth. Inflorescence a single terminal many-flowered terete spike-let. Glumes many, imbricate all round the rhachis; the lowest 1 or 2 empty, shorter than the spikelet; many succeeding ones hermaphrodite and fruit-bearing; the uppermost male or sterile. Hypogynous bristles usually 6, but varying from 3 to 8, rarely absent. Stamens 3 or fewer. Style swollen at the base; branches 3 or 2, linear. Nut obovoid, trigonous or plano convex.

Species estimated at 115, distributed. over the whole world, but most numerous in America. Of the 5 found in New Zealand, 2 are endemic, 2 extend to Australia, the remaining one is almost cosmopolitan.

A.Limnochloa. Stem stout, spikelet large, hardly wider than the stem. Glumes subrigid.

Stems stout, septate. Spikelet 1–2 in. long 1. E. sphacelata.

B.Eleogenus. Stem slender. Spikelets small, broader than the stem. Glumes membranous. Style 2-fid.

Seems short, in. 1–2½ Spikelet ⅙–¼ in. 2. E. neo-zealandica.

C. Eu-Eleocharis. Stem slender. Spikelet small, broader than the stem Glumes membranous. Style 3-fid.

Stems 2–6 in., filiform. Spikelets ⅛–⅕ in, compressed. Nut longitudinally ribbed and transversely striate 3. E. acicularis.
Stems 4–18 in., rather stout. Leaf-sheath truncate with an erect mucro. Spikelec ¼–¾ in. Nut smooth 4. E. acuta.
Stems 3–15 in., very slender. Leaf-sheath oblique, acute. Spikelet ⅛–¾ in. Nut smooch 5. E. Cunning hamii
1.E. sphacelata, R. Br. Prodr. 224.—Rhizome stout, creeping, stolon if erous. Stems stout, cylindrical, 1–3 ft. high, ⅓ in. diam., hollow, transversely septate; sheaths long, membranous. Spikelet very large, 1–2 in. long, ⅓ in. diam., solitary, terminal, cylindrical, pale-coloured, tip acute. Glumes numerous, very closely imbricate, obovate-oblong, obtuse, 1-nerved, membranous, pale with a brown line just inside the scarious margin. Hypogynous bristles 6–9, usually exceeding the nut, retrorsely scabrid. Style very long, branches 3. Nut broadly obovoid, compressed, minutely granular, pale, crowned with the persistent dark-brown conic swollen base of the stvle.—A. Cunn. Precur. n. 277; Raoul, Choix, 40; Hook. f. Fl. Nov. Zel. i. 269; Handb. N.Z. Fl. 300; Benth. Fl. Austral. vii. 292 (Heleocharis). page 768

North Island: Wet swamps and margins of lakes, not uncommon South Island: Nelson—Takaka, Kirk! Canterbury, Armstrong. Westland—Okarito, A. Hamilton! Southland—Bluff Island, Lyall. Stewart Island: Head of Paterson's Inlet, Petrie! Sea-level to 1500 ft. December–February.

An abundant Australian and Tasmanian plant, and very closely allied to the widely diffused E. plantaginea, R. Br.

2.E. neo-zealandica, C. B. Clarke ex T. Kirk in Trans. N.Z. Inst. xxvi. (1894) 260.—Rhizome slender, creeping, 1–3 in. long. Stems shore, 1–2½ in. high, striate; sheath membranous, mouth oblique. Spikelet solitary, terminal, ⅙–¼ in. long, broadly ovoid, much wider than the stem, 4–8-flowered. Glumes broadly ovate, concave, obtuse, red-brown with a paler centre and searious margins. Hypogynous bristles wanting. Stamens 3. Style with 2 linear arms and a very small swollen base. Nut obovoid, biconvex, smooth, pale-brown.

North Island: Auckland—Sand-dunes between Cape Maria van Diemen and Ahipara, T. F. C. South Island: Nelson—Cape Farewell, Kirk!

A curious little species, belonging to the section Eleogenus, characterized by the spikelets broader than the slender stems, membranous glumes, and 2-fid style. Mr. Clarke remarks that it approaches the section Isolepis of Scirpus in the absence of hypogynous bristles and the reduced size of the swollen base of the style.

3.E. acicularis, R. Br. Prodr. 224.—Rhizome very slender, almost filiform, creeping, stoloniferous. Stems numerous, tufted, extremely slender, capillary, 2–6 in. high; sheaths membranous, acute. Spikelet small, slender, ⅛–⅕ in. long, compressed, pale to dark-brown, 3–6-flowered. Glumes ovate, obtuse, membranous, brown with a greenish or pale centre and narrow searious margins. Hypogynous bristles 2–4, short, deciduous. Style-branches 3; style-base small, conic, depressed. Nut small, pale, obovoid-oblong, longitudinally ribbed and with minute transverse striæ between the ribs.—Kunth, Enum. ii. 141; C. B. Clarke in Hook. f. Fl. Brit. Ind. vi. 628.

South Island: Otago—Lake Te Anau, Petrie! (No. 1647).

I have seen no specimens but Mr. Petrie's, which are in young flower only. Mr. C. B. Clarke, who has examined them, states that he is satisfied that they belong to the small group consisting of E. acicularis and a few very closely allied species, and most probably to E. acicularis itself, which is an almost cosmopolitan plant, although not yet recorded from Australia.

4.E. acuta, R. Br. Prodr. 224.—Rhizome creeping. Stems numerous, tufted, 4–18 in. high, rather stout or slender, striate; sheath closely appressed to the stem, with a horizontally truncate mouth, the margin of which is thickened and usually dark-coloured, with a small erect mucro or rudimentary lamina on one side. page 769Spikelet variable in length, ¼–¾ in. or more, linear-oblong, cylindric, obtuse or subacute, many-flowered. Glumes broadly ovate, obtuse, membranous, concave, brown with usually a pale line down the centre; margins broad, scarious. Hypogynous scales 4–8, exceeding the nut. Stamens 3. Style-branches 3. Nut broadly obovoid, biconvex, smooth or very minutely pitted, brown, crowned by the small conic style-base.—Hook. f. Handb. N.Z. Fl. 745; Benth. Fl. Austral. vii. 294 (Heleocharis). E. gracilis (excl. var. b and y), Hook. f. Fl. Nov. Zel. i. 270; Handb. N.Z. Fl. 301 (not of R. Br.). E. ambigua, Kirk ex Buch. in Trans. N.Z. Inst. vi. (1874) 225.

North and South Islands, Chatham Islands: Abundant in wet places from the North Cape to Foveaux Strait. Sea-level to 2000 ft. November–March.

Also plentiful in Australia, Tasmania, and Norfolk Island.

5.

E. Cunninghamii, Boeck. in Flora, xli. (1858) 412 (Heleocharis).—Rhizome long, creeping, scaly, stout or slender, dark red-brown or almost black. Stems many from the rhizome, variable in length, 3–15 in., very slender, sometimes almost filiform, striate; sheath membranous, with a thin oblique mouth. Spikelet small, short, ⅛–¼ in. long, broadly ovoid, subacute, 5–20-flowered. Glumes broadly oblong, obtuse, membranous, usually pale but often stained with red-brown; margins broad, scarious. Hypogynous bristles 4–8, exceeding the nut. Style-branches 3. Nut broadly obovoid, plano-convex or obscurely trigonous, smooth, pale yellowish-brown, crowned by the small pyramidal style-base, which is rather broader than the top of the nut.—Linnœa, xxxvi. (1889–70) 427. E. gracilis, var. gracillima and var. radicans, Hook. f. Fl. Nov. Zel. i. 270; Handb. N.Z. Fl. 301 (not of R. Br.). E. gracillima, Hook. f. Handb. N.Z. Fl. 745. Heleocharis Hookeri, Boeck. in Linnæa, xxxvi. (1869–70) 430. Isolepis acicularis, A. Rich. Fl. Nouv. Zel. 103 (not of R. Br.).

North and South Islands.—Wet places from the North Cape to Foveaux Strait, but often local in the South Island. Sea-level to 2000 ft. November–March.

Hooker's two varieties gracillima and radicans, originally published as forms of the Australian E. gracilis, only differ in size and degree of development, radicans being clearly a depauperated state. In many localities they can be seen to merge into one another. Boeckeler, probably working upon scanty herbarium material, treated them as distinct species, publishing his E. Cunninghamii, which answers to var. radicans, in 1858, and E. Hookeri, which is equivalent to var. gracillima, in 1869. E. Cunninghamii has several years' priority over Hooker's gracillima, which was not published until 1867, but it is unfortunate that it was not originally applied to what must be considered the type of the species, Hooker's var. gracillima.