M. australis, Meissn. Gen. Comm. 227.—A much-branched climber, covering shrubs or small trees, or trailing over cliffs or rocks; trunk stout, woody, sometimes 3 in. diam.; bark greyish; branches numerous, flexuous and interlaced, the younger ones grooved and angled, glabrous or faintly scaberulous. Leaves petio-late, 1–3 in. long, ovate or orbicular-oblong, apiculate or acuminate, rarely obtuse, cordate or truncate at the base, thin and membranous, quite glabrous, usually entire, but sometimes panduriform or 3-lobed, margins undulate-crisped; petioles ⅓–1 in. long; stipules deciduous. Panicles large, axillary and terminal, much branched, 1½–3 in. long or more. Flowers about ⅙ in. diam., greenish, diœcious; males with 8 stamens and the rudiment of an ovary; females with 8 blunt staminodia and a short triquetrous ovary; stigmas broad, fimbriate. Fruiting-perianth closed over the fruit, slightly succulent or almost herbaceous. Nut black, shining, 3-angled and 3-grooved, angles often twisted.—M. adpressa, Hook. f. Handb. N.Z. Fl. 236 (not of Meissn). Polygonum australe, A. Rich. Fl. Nouv. Zel. 178; A. Cunn. Precur. n. 355; Raoul, Choix, 42. P. adpres-sum, A. Cunn. I.c. n. 356 (not of Labill.). Coccoloba australis, Forst. Prodr. n. 176.

North and South Islands, Stewart Island, Chatham Islands: Common from the Three Kings Islands and the North Cape southwards. Sea-level to 2000 ft. November–April.

Also found in Norfolk Island, and very closely allied to the Australian M. adpressa, Meissn.

M. complexa, Meissn. Gen. Comm. 227. — Forming dense thick and elastic prostrate masses many feet in diam., or climbing over bushes or rocks. Stems slender, tough and woody, much interlaced; branches very numerous, flexuous, terete, the ultimate ones pubescent with short stiff hairs; bark dark red-brown. page 593Leaves petiolate, exceedingly variable in size and shape, even on the same plant, ⅕–¾ in. long or more, broadly oblong or obovate or orbicular, rounded or retuse at the tip, rarely subacute, cordate or truncate or rounded at the base, often contracted in the middle, sometimes 3-lobed, quite smooth, glabrous or the petiole puberulous, coriaceous or almost membranous, margins even, entire; stipules deciduous. Spikes axillary and terminal, long or short, simple or compound, glabrous or puberulous, sometimes reduced to 2 or 3 flowers, at other times forming a panicle 1–1½ in. long or more. Flowers small, diœcious. Stamens 8. Stigmas very broad, fimbriate. Fruiting-perianth much enlarged, often ¼–⅓ in. diam., succulent, waxy-white, forming a fleshy cup surrounding the fruit, in some varieties unaltered and herbaceous. Nut black, shining, deeply triquetrous.—Hook. f. Handb. N.Z. Fl. 236. M. microphylla, Col. in Trans. N.Z. Inst. xx. (1888) 204. M. paucifolia, M. trilobata, and M. truncata, Col. I.c. xxi. (1889) 99–101. Polygonum com-plexum, A. Cunn. Precur. n. 357; Raoul, Ghoix, 42; Hook. f. Fl. Nov. Zel. i. 210.

North and South Islands: Abundant throughout, ascending to 2000 ft. Pohuehue. November–April.

Varying much in the size and shape of the leaves, the degree of development of the spikes, and the extent to which the perianth enlarges and becomes fleshy in fruit. Mr. Colenso made no less than 4 species, based mainly on variations in the above characters, but his own specimens show how inconstant these distinctions are, and how little they can be relied upon.

M. axillaris, Walp. Ann. i. 552.—A small much-branched prostrate or diffuse shrubby plant, usually forming densely matted patches 3–12 in. diam., but sometimes open and straggling; stems and branches woody; branchlets puberulous. Leaves on rather long petioles, small, 1/10–⅓ in. long, broadly oblong or ovate-oblong or almost orbicular, obtuse or retuse, rounded at the base, flat, quite glabrous, dotted beneath. Flowers small, solitary or 2. together in the axils of the leaves, or the males rarely forming short few-flowered spikes at the ends of the branchlets; pedicels slender. Stigmas fimbriate. Perianth succulent in fruit or almost unaltered. Nut triquetrous with obtuse angles, black, smooth and shining, rather longer than the perianth.—Hook. f. Handb. N.Z. Fl. 236; Benth. Fl. Austral. v. 275. M. hypogæa, Col. in Trans. N.Z. Inst. xxi. (1889) 98. Polygonum axillare, Hook. f. Fl. Nov. Zel. i. 211.

North Island: Mountain districts in the interior, from Taupo and the East Cape southwards. South Island: Common in mountain districts throughout. Sea-level to 4500 ft. December–March.

Also found in Tasmania and Australia, and reported from Lord Howe Island.

M. ephedrioides, Hook. f. Fl. Nov. Zel. i. 211.—A much-branched prostrate shrub; stems 9–36 in. long or more, rigid and wiry, deeply grooved, leafless and rush-like or sparingly leafy; page 594branchlets often scaberulous. Leaves when present few and scattered, periolate or almost sessile, ⅙–1 in. long, linear or linear-lanceo-late, often dilated or almost hastate at the base, acute or obtuse, glabrous, sometimes scaberulous on the midrib beneath; stipules short, obliquely truncate. Flowers small, polygamous; those on the male plant in lax axillary simple or branched glabrous spikes, often with a few female flowers intermixed; on the female plant in few-flowered fascicles or short dense spikes, usually with 1 or two male flowers mixed with the females. Stigmas fimbriate. Perianth succulent or remaining unaltered in fruit. Nut exceeding the perianth, black, smooth and shining, triquetrous with the angles obtuse. —Handb. N.Z. Fl. 237.

Var. muricatula.—Smaller and much more slender, branchlets often almost filiform. Leaves usually present, ⅙–½ in. long, linear. Perianth-segments membranous in fruit.—M. muricatula, Col. in Trans. N.Z. Inst. xxii. (1890) 482 (perhaps a distinct species).

North Island: Hawke's Bay—Near the sea, Colenso! A. Hamilton! Upper Rangitikei, Bishop Williams, Petrie! South Island: Marlborough — Near Blenheim, Buchanan! Kirk! Canterbury—Waipara, Haast! Otago—Lower Waitaki, Hector and Buchanan; Kurow, Awamoko, Roxburgh, Petrie! Var. muricatula: North Island—Lake Taupo, Tryon! Ruapehu, H. Hill! Waipawa, A. Hamilton! Sea-level to 3000 ft. December–March.

Easily recognised by the prostrate rush-like stems, which are often entirely leafless.