Other formats

    TEI XML file   ePub eBook file  

Connect

    mail icontwitter iconBlogspot iconrss icon

Manual of the New Zealand Flora.

16. Cotula, Tourn

16. Cotula, Tourn.

Creeping or tufted perennial or annual herbs, usually of small size, often aromatic. Leaves alternate, pinnatifid or pinnatisect, rarely entire or toothed. Heads small, peduncled, heterogamous and discoid or rarely homogamous through the suppression of the female florets, sometimes dioecious. Involucre hemispheric or campanulate; bracts in about 2 series, membranous or herbaceous; margins often scarious. Receptacle flat or convex or conical, page 350without scales. Female florets exterior, in 1 or 2 series, fertile; corolla broad or conic or wanting. Disc-florets hermaphrodite or often male; corolla regular, tube slender or stout and 2-winged, limb 4-toothed. Anthers obtuse at the base, entire. Style-branches of the disc-florets truncate or obtuse, sometimes undivided. Achenes compressed, sometimes winged. Pappus wanting.

A genus of 50 or 60 species, scattered widely over the world in both temperate and tropical regions. Several of the New Zealand species are difficult of discrimination, and require further study with more copious suites of specimens.

A. Eucotula. Receptacle flat or convex. Female florets without any corolla.

Achenes of the female florets stipitate, in a single row. Stout, glabrous, 2–10 in. high. Leaves lanceolate, ½–2 in., variously toothed or lobed. Heads yellow,⅓–½ in. diam 1. C. coronopifolia.

B. Strongylosperma. Receptacle flat or convex. Female florets without any

corolla. Achenes of the female florets in several rows.

Slender, diffuse, silky. Leaves pinnate or bipinnate. Heads small, 1/10–⅙ in. diam 2. C. australis.

C. Leptinella. Receptacle convex or conical. Female florets in 1 or several series; corolla always present, usually inflated at the base.

* Heads bisexual.

Stout, fleshy. Leaves ½–1 in., much divided. Peduncles leafy. Heads black or dark-brown. Florets tubular or cylindric, rugose 3. C. atrata.
Stout, softly woolly. Leaves 2–6 in., 3–4-pinnatisect. Heads ⅓–½ in. diam.; florets eglandular 4. C. plumosa.
Stout, woolly. Leaves rather fleshy, 1–3 in., pinnate or pinnatifid. Heads ¼–⅓ in.; florets glandular 5. C. lanata.
Stout, silky. Leaves 2–5 in., pinnatifid. Heads ⅓ in.; florets eglandular, females in many rows 6. C. Muelleri.
Slender, silky or glabrate. Leaves membranous, 1–3 in., pinnatifid. Heads ⅓ in.; florets glandular, females in 1 row 7. C. Traillii.
Small, silky, densely matted; stems ½–3; in. Leaves ⅙–⅓ in., pinnatifid. Heads almost sessile, 1/10–⅛ in. diam.; female florets in 1–2 series 8. C. Maniototo.
Stems long, slender, creeping. Leaves membranous, ½–2; in., pinnatifid; segments broad, deeply toothed. Peduncles longer or shorter than the leaves. Heads ⅙–⅕ in. diam.; female florets numerous, in 3–4 series 9. C. minor.
Stems slender, wiry. Leaves ¼ in., pinnatifid. Heads: 1/10 in. diam. 10. C. filiformis.
Stems rather stout. Leaves pubescent, ½–1½ in., pinnatifid; segments narrow, incised. Peduncles much longer than the leaves. Heads ⅙–¼ in.; female florets in 3–4 series 11. C. Haastii.
Stems stout, wiry. Leaves rigid, pectinately pinnatifid; segments entire. Peduncles much longer than the leaves. Heads⅕–¼ in.; female florets in 3–4 series 12. C. pectinata.
Minute, tufted. Leaves imbricate, ⅙–⅓ in., cut straight down from the tip into 5–7 subulate lobes. Heads small, nearly sessile, ⅛ in. diam. 13. C. Goyeni.page 351
Stems stout, 6–12 in. long. Leaves ¾–2 in., obovate-spathulate, crenately 3–5-toothed at the tip. Peduncles short. Heads in. diam.; female florets in many-series 14. C. Featherstonii.

** Heads unisexual.

Leaves ½–1½ in., linear-spathulate, quite entire. Peduncles 2–4 in., bracteate. Heads ¼–⅓ in. daim. 15. C. linearifolia.
Leaves ½–1½ in., pinnatifidly cut into 5–8 linear entire segments. Peduncles 1–4 in., bracteate. Heads ⅓–¾ in. diam. 16. C. pyrethrifolia.
Slender, wiry, silky. Leaves ¼–1 in., silky on both surfaces, deeply pinnatifid. Peduncles short, stout, ebrac-teate. Heads ⅛–¼ in.; females the largest 17. C. perpusilla.
Slender, creeping, 4–12 in. long. Leaves membranous, 1–2 in., deeply pinnatifid, segments incised. Peduncles 1–3 in., ebracteate. Heads ⅙–⅓ in.; females the largest 18. C. squalida.
Stout or slender. Leaves ½–2 in., obovate or spathulate, crenate or lobulate, pinnatifid below. Peduncles longer or shorter than the leaves. Heads ⅙–⅓ in.; females the largest 19. C. dioica.
1.C. coronopifolia, Linn. Sp. Plant. 892.—Perfectly glabrous smooth and succulent. Stems creeping and rooting below, ascending at the tips, branched, 2–10 in. high. Leaves scattered, dilated and clasping the stem at the base, ½–2 in. long, linear-lanceolate or oblong-lanceolate, coarsely toothed or lobed or pinnatifid or entire, Heads bright - yellow, ⅓–½ in. diam., on long slender axillary peduncles exceeding the leaves; involucral bracts linear-oblong, obtuse, membranous. Receptacle slightly convex. Female florets in 1 series, on slender flattened pedicels; corolla wanting; ovary winged, notched at the top; style short, seated in the notch. Discflorets on much shorter pedicels, very numerous; corolla tubular, more or less dilated at the base, 4-toothed at the tip. Achenes of the female florets oblong, compressed, with a broad spongy wing; those of the disc-florets smaller, with a much narrower wing.— Forst. Prodr. n. 300; A. Rich. Fl. Nouv. Zel. 235; A. Cunn. Precur. n. 443; Raoul, Choix, 45; Hook. f. Fl. Nov. Zel. i. 127; Handb. N.Z. Fl. 141; Benth. Fl. Austral. iii. 549; Kirk, Students' Fl. 322.

Var. integrifolia, Kirk, l.c.—Stems small, 1–2 in., almost simple. Leaves linear, entire, obtuse. Peduncle terminal, slender. Head small, in. diam.— C. integrifolia, Hook. f. Fl. Tasm. i. 192, t. 50B.

North and South Islands, Stewart Island, Chatham Island: Not uncommon in wet places in lowland districts. October–February.

Widely spread through the south temperate zone; also found in some parts of Europe and North America, but probably naturalised only.

2.C. australis, Hook. f. Fl. Nov. Zel. i. 128.—A slender much-branched flaccid herb 2–6 in. high; branches spreading, prostrate or suberect, more or less clothed with long lax hairs or almost page 352glabrous. Leaves ½–1 in. long, oblong in outline, membranous, deeply pinnatifid or bipinnatifid; ultimate segments linear, entire, acute or mucronate. Heads 1/10–⅕ in. diam., on long slender peduncles; involueral bracts in 2 series, linear-oblong, obtuse. Female florets numerous, in 3 series, on slender pedicels; corolla wanting. Disc-florets comparatively few, subsessile; corolla tubular, dilated at the base, 4-toothed at the tip. Achenes of the female florets obovate, with a broad thick wing, glandular on both faces; those of the disc-florets much smaller, glabrous, hardly winged.— Handb. N.Z. Fl. 141; Fl. Tasm. i. 191, t. 50A; Benth. Fl. Austral. iii. 550; Kirk, Students' Fl. 322. G. venosa, Col. in Trans. N.Z. Inst. xxiii. (1891) 388. Strongylosperma australe, Less. Syn. Comp. 261.

Kermadec Islands, North and South Islands, Chatham Islands: Abundant in lowland districts. September–March.

Also found in Australia, Tasmania, and Tristan d'Acunha; and perhaps not really different from a South African plant.

3.C. atrata, Hook. f. Handb. N.Z. Fl. 142.—Rhizome stout, creeping, tortuous; roots long and stringy. Stems 1–6 in. high, erect or ascending, stout, leafy, densely pubescent. Leaves ½–1½ in. long, thick and fleshy, pubescent on both surfaces; petiole broad, flat, sheathing at the base; blade linear-oblong or linear-obovate in outline, pinnatifid; segments close-set, entire or toothed or again pinnatifid; cauline leaves smaller, pinnatifid or lobed. Heads subglobose, ⅓–¾ in. diam., black or brownish-yellow; involucral bracts in 2–4 series, linear-obovate, entire or pinnatifid, shorter than the florets or equalling them. Female florets in 3–5 series; corolla cylindric, rugose, 3–4-toothed. Disc-florets with a long rugose tube and narrow funnel-shaped 4-tootbed limb. Achenes linear-oblong or linear-obovoid, rugose.—Kirk, Students' Fl. 323.

South Island: Dry shingle slopes in alpine localities, Nelson to Otago; not uncommon, 3500–6500 ft. January–February.

Easily recognised by the greyish-green foliage and almost black heads. It is a variable plant in the size of the heads, length of the involucral bracts, and in their being sometimes entire, sometimes pinnatifid.

4.C. plumosa, Hook. f. Handb. N.Z. Fl. 141.—A large densely tufted aromatic species, often forming broad soft patches, every where clothed with long villous hairs. Stems short, stout, creeping. Leaves on long slender petioles 3–6 in. long; blade 2–6 in., oblong in outline, flaccid and membranous, finely 3–4-pinnatifid; primary divisions close-set, linear-oblong, recurved, 2-pinnatifid on the upper side; ultimate segments ⅙ in. long, linear, toothed on one side. Peduncles slender, shorter than the leaves, usually with a linear entire or pinnatifid bract about the middle. Head ⅓–½ in. diam.; involucral bracts in 2 or 3 series, broadly oblong, with broad page 353purplish-black margins. Receptacle conical. Female florets in 2–3 series, shortly pedicelled; corolla compressed, swollen at the base, contracted towards the mouth, unequally 4-toothed. Discflorets funnel-shaped, 5-toothed. Achenes of the female florets obovoid; those of the disc-florets minute, abortive.—Kirk, Students' Fl. 323. Leptinella plumosa, Hook. f. Fl. Antarct. i. 26, t. 20.

Auckland and Campbell Islands: Hooker! Kirk! Chapman! Antipodes Island: Kirk! Macquarie Island: Scott, A. Hamilton, December–January.

A very handsome species, easily recognised by the large pale-green feathery foliage. It is also found in Kerguelen Island and the Crozets.

5.C. lanata, Hook. f. Handb. N.Z. Fl. 141.—Stems stout, prostrate and rooting below, ascending at the tips, branched, 3–12 in. long, densely clothed with white woolly hairs or glabrate. Leaves 1–3 in. long, woolly or glabrate, rather thick, with broad sheathing petioles; blade oblong in outline, pinnate or pinnatifid; pinnæ close-set, curved, 3–5-toothed or -lobed along the upper edge, minutely glandular. Peduncles terminal, shorter than the leaves, stout, woolly. Heads ¼–⅓ in. diam.; involucral bracts in several series, broadly oblong, obtuse, green. Florets all studded with pellucid conglobate glands; females ovate-oblong, broadest at the base, narrow at the mouth and minutely 4-toothed; disc-florets narrow funnel-shaped, 5-toothed. Achenes of the female florets obovate, compressed; those of the disc-florets minute, abortive. — Kirk, Students' Fl. 323. Leptinella lanata, Hook. f. Fl. Antarct. i. 25, t. 19. L. propinqua, Hook. f. l.c. 27.

Auckland and Campbell Islands: Hooker, Kirk! Chapman! December–January.

Distinguished from the preceding by the smaller size, stouter habit, leaves not so finely cut, smaller heads, and glandular florets.

6.C. Muelleri, T. Kirk, Students' Fl. 324.—Stems long, rather stout, creeping and rooting, branched, sparingly villous towards the tips; branches ascending. Leaves 2–5 in. long; petiole ¾–2 in., sheathing at the base; blade 1–3 in., linear-obovate, membranous, glabrate or slightly silky, gland-dotted, deeply pinnatifid; segments oblong, closely toothed or lobed, teeth acute. Peduncles 1–3 in. long, usually shorter than the leaves, pubescent. Heads ⅓ in. diam.; involucral bracts in about 3 series, oblong-ovate, obtuse or subacute, membranous. Female florets in many rows, usually more numerous than the disc-florets; corolla short, ovoid, not glandular. Disc-florets larger; corolla tubular, 4-toothed; style undivided. Achenes of the female florets clavate, tetragonous.— Leptinella potentillina, F. Muell. Veg. Chath. Is. 28, t. 6. page 354

Chatham Islands: Not uncommon in moist places, H. H. Travers! Enys! Cockayne and Cox!

This was apparently referred to C. lanata by Sir J. D. Hooker (Handb., p. 733), but it differs from that plant in being much less woolly, in the larger leaves with broader toothed segments, longer peduncles, and glandless florets.

7.C. Traillii, T. Kirk, Students' Fl. 324.—Stems slender, creeping and rooting, 3–12 in. long, silky towards the tips. Leaves 1–3 in.; petiole slender, sheathing at the base; blade linear-obo- vate, membranous, glabrate or with scattered lax hairs on both surfaces, deeply pinnatifid; segments close-set, broadly oblong, toothed on the upper margin and round the apex, teeth apicu- late or hair-tipped. Peduncles axillary, pubescent, usually shorter than the leaves. Heads ⅓ in. diam., involucral bracts in 2–3 series, orbicular-ovate, membranous, with scarious purplish margins. Female florets few, in 1 series, shortly stipitate; corolla ovoid, compressed. Disc-florets very numerous, funnel-shaped, 4-lobed; style undivided, with a discoid stigma. Achenes of the female florets broadly ovoid or almost orbicular, turgid, 3-winged.

Stewart Island: Chiefly on sand-dunes, Kirk! December–February.

Very closely allied to C. Muelleri, of which it may be a variety. The habit is the same, and the leaves can only be distinguished by the smaller size and apiculate segments. But the heads are markedly different in the few female florets, which are in one row only, and have much broader achenes.

8.C. Maniototo, Petrie in Trans. N.Z. Inst. xiv. (1882) 362.— Small, densely matted, forming broad greyish patches; everywhere covered with long silky hairs. Stems ½–3 in. long, creeping and rooting; branches very short, leafy. Leaves numerous, silky on both surfaces, dilated and sheathing at the base, ⅙–⅓ in. long, linear- oblong, deeply pinnatifid; segments narrow-linear, acute. Heads terminating the branches, almost sessile, minute, 1/10–⅛ in. diam.; involucral bracts in 2 series, broadly oblong or obovate, obtuse, silky on the outside, membranous. Female florets in 1 or 2 series; corolla narrow-tubular, almost filiform, with a 2-lipped mouth; style exserted, 2-fid. Disc-florets much larger; corolla funnel- shaped with a long tube, 4–5-toothed; anthers and style exserted. Achenes of the female florets oblong, smooth and turgid, hardly winged; those of the disc-florets minute, abortive.—Kirk, Students' Fl. 323.

South Island: Canterbury—Lake Lyndon, Enys! Kirk! T. F. C.; Lake Tekapo, T. F. C. Otago—Kakanui, Maniototo Plain, Nevis Valley, Moss-burn, Lake Te Anau, Petrie! Buchanan! Sea-level to 3000 ft. January–February.

A very distinct little plant, probably common throughout the South Island.

9.

C. minor, Hook. f. Handb. N.Z. Fl. 142.—Stems slender, creeping, branched, 2–12 in. long or more, glabrous or silky at the page 355 tips. Leaves alternate on slender creeping runners or fascicled on short lateral branchlets, thin and membranous, glabrous or slightly silky, ½–2 in. long, linear-oblong or linear-obovate in outline, pin-natifid almost to the base; segments close-set or the lower distant, oblong or obovoid, recurved, deeply and acutely toothed on both edges or on the upper only. Peduncles slender, naked, shorter or longer than the leaves. Heads small, ⅙–⅕ in. diam.; involucral bracts few, 8–12, broadly oblong or orbicular, usually with broad purplish margins. Female florets in 3–4 series; corolla inflated, broadly ovoid, with a narrow 2–3-toothed mouth. Disc-florets fewer in number, funnel-shaped, 4-toothed. Achenes of the female florets obovoid, glandular.—Kirk, Students' Fl. 324. Leptinella minor, Hook. f. Fl. Nov. Zel. i. 129. Soliva tenella, A. Cunn. Precur. n. 445.

North and South Islands: Not uncommon from the North Cape southwards. Sea-level to 2500 ft. November–January.

10. C. filiformis, Hook. f. Handb. N.Z. Fl. 142. —"A very slender rigid creeping plant, glabrous or pilose. Leaves minute, ¼ in. long, oblong, pinnatifid, segments subulate. Scapes filiform, 1 in. long, naked. Heads minute, 1/10 in. diam.; involucral scales 6–8, orbicular, with purple edges; receptacle conical; ray-florets about 20; corolla short, compressed, inflated, very broad-oblong, 2-lobed above; achene obconic, glandular; disc-florets funnel- shaped, 4-lobed; lobes glandular."—Kirk, Students' Fl. 325.

South Island: "Canterbury Plains, amongst grass, Haast."

This does not seem to have been collected since its original discovery, more than forty years ago. Not having seen specimens, I have reproduced Hooker's description.

11.C. Haastii, T. Kirk, Students Fl. 325.—Stems rather short, stout, creeping, branched, 1–6 in. long or more, pubescent. Leaves ½–1½ in. long, pubescent on both surfaces, gland-dotted, linear- oblong or linear-obovate, deeply pinnatifid or almost pinnate; segments narrow-oblong, recurved, entire or deeply toothed or incised, teeth often confined to the upper margin. Peduncles slender, pubescent, much longer than the leaves. Heads ⅙–¼ in. diam.; involucral bracts in 2–3 series, broadly oblong, obtuse, with broad purple tips. Receptacle convex. Female florets in 3–4 series; corolla short, ovoid, compressed. Disc-florets very numerous, funnel-shaped, with 4 short and broad teeth. Achene oblong-obovoid, obscurely angled, glandular.

South Island: Canterbury Plains, Haast! Petrie; Banks Peninsula, Haast! Kirk! Cockayne! Sea-level to 2500 ft. December–January.

Closely allied to C. pectinata, but the leaves are broader, with the segments deeply toothed or incised.

page 356
12.C. pectinata, Hook. f. Handb. N.Z. Fl. 142.—Stems 1–6 in. long, creeping and rooting, rather stout, wiry, woolly or glabrate, often putting out silky runners. Leaves few, scattered, rigid, glabrous or silky, ¾–1½ in. long, linear-oblong in outline, pectinately pinnatifid; segments short, subulate, entire. Peduncles slender, naked or rarely with a minute bract about the middle, 1–3 in. long, exceeding the leaves. Heads ⅕–¼ in. diam., involucral bracts in 2–3 series, broadly oblong, pubescent, purplish; margins jagged. Female florets in several series; corolla ovoid, compressed, minutely 2–4-toothed at the narrow mouth. Disc-florets funnel-shaped, 4-toothed. Achene narrow-obovoid, compressed and slightly winged. —Kirk, Students' Fl. 325.

Var. sericea, Kirk, l.c. 326.—Smaller and more compactly branched, everywhere clothed with long dense silky hairs. Peduncles shorter and stouter, ½–1 in. long. Heads in. diam. Perhaps a distinct species.

South Island: Nelson—Clarence Valley, T. F. C.; Mount Captain, Kirk! Canterbury—Mount Torlesse, Haast! mountains above the Broken River, Enys! Kirk! T. F. C. Oago—Lake district, Hector and Buchanan! common in mountain districts, Petrie! Var. sericea: Old Man Range and Mount Cardrona, Petrie! Ben Lomond, Cockayne! Altitudinal range from 1500 to 6000 ft. December–January.

13.C. Goyeni, Petrie in Trans. N.Z. Inst. xviii. (1886) 295.— A small tufted species. Stems 1–3 in. long, creeping and rooting, much branched; branches short, ascending at the tips, with the leaves ¼ in. diam. Leaves alternate, imbricate, appressed to the branch, ⅛–⅕ in. long; lower half broad, membranous, glabrous or more or less pubescent, sheathing at the base; upper half cut straight down into 5–7 linear-subulate erect lobes. Heads terminal, minute, ⅛ in. diam., on short woolly peduncles rarely exceeding the leaves; involucral bracts in 1 or 2 series, ovate-oblong, with scarious purple margins. Female florets few, in 1 series; corolla ovoid, compressed, narrowed at the mouth. Disc-florets numerous, funnel-shaped. Mature achenes not seen.—Kirk, Students' Fl. 326.

Var. pinnatisecta, Kirk, l.c.—Leaves pinnatifid, clothed with silky hairs.

South Island: Otago—Mount Pisa and the Hector Mountains, Petrie! 5000–6000 ft. January–February.

A curious little plant, perhaps nearest to C. pectinata. The leaves closely resemble those of Azorella Selago.

14.C. Featherstonii, F. Muell. ex Hook. f. Handb. N.Z. Fl. 733.—Stems 6–12 in. long or more, rather stout, prostrate or decumbent at the base, ascending above, much branched, leafy, clothed with short soft pubescence. Leaves alternate, ¾–2 in. long, obovate-spathulate or oblong-spathulate, gradually narrowed to a sessile base, crenately 3- or 5-toothed at the tip or entire, flat, fleshy, finely and softly pubescent on both surfaces. Peduncles page 357 axillary and terminal, ½–1 in. long. Heads ⅓ in. diam., yellow; involncral bracts 10–15, in about 2 series; outer ovate-lanceolate, inner broadly oblong. Female florets in many series; corolla ovoid or conical, swollen at the base, obscurely toothed at the contracted mouth. Disc-florets numerous, tubular, 4-toothed. Achenes narrow - obovoid, prominently costate, glandular.—Kirk, Students Fl. 326. Leptinella Featherstonii, F. Muell Veg. Chath. Is. 27, t. 5.

Chatham Island: Usually near the coast, H. H. Travers! Cockayne and Cox!

A very remarkable species, quite unlike any other.

15. C. linearifolia, Cheesem. in Trans. N.Z. Inst. xv. (1883) 299.—Small, dark-green, rather thick and fleshy, aromatic, spar ingly pilose. Stems prostrate, branched, ascending at the tips. Leaves alternate, ½–1½ in. long, narrow-linear or linear-spathulate, obtuse, gradually narrowed to the sheathing base, quite entire, coriaceous, gland-dotted. Peduncles 2–4 in. long, slender, ter minal, with 4–8 small linear bracts. Heads unisexual, ¼–⅓ in. diam.; involucral bracts in about 3 series, linear-oblong, obtuse, with purplish scabious margins. Receptacle convex. Florets studded with transparent glands; female corolla swollen at the base, obscurely tetragonous, narrowed above, minutely 4-toothed; corolla of the males smaller and more slender, narrow funnel- shaped, 4-lobed. Aohene linear-obovoid, compressed.—Kirk, Students' Fl. 326.

South Island: Nelson—Mountains flanking the Wairau Valley, T. F. C. 3000–4500 ft. December–January.

Distinguished from C. pyrethrifolia by the entire leaves. In outward appearance it closely resembles Ahrotanella linearis.

16.C. pyrethrifolia, Hook. f. Handb. N.Z. FL 142.—Stout, rather fleshy, aromatic, glabrous or sparingly pilose. Rhizome creeping, often branched; stems prostrate or decumbent at the base, ascending at the tips. Leaves alternate, fleshy or coriaceous, ½–1½ in. long; petiole usually half the length, sheathing at the base; blade pinnatifidly cut into 5–8 alternate narrow-linear to linear-oblong obtuse segments 1/10–¼ in. long. Peduncles variable in length, ½–4 in., with 1 or several linear bracts. Heads unisexual, large, ⅓–¾ in. diam.; involucral bracts in 2–3 series, linear-oblong, obtuse, herbaceous, with purplish scarious margins. Receptacle convex. Florets glandular; corolla of the females inflated and truncate at the base, 4-toothed at the mouth; of the males funnel-shaped, 4-lobed. Achene narrow-obovoid.—Kirk, Students' Fl. 327.

South Island: Abundant in mountain districts in Nelson, Marlborough, and Canterbury, rare and local in Otago. 2500–6000 ft. December- February.

page 358
17.C. perpusilla, Hook. f. Handb. N.Z. Fl. 143.—Often forming broad patches, more or less silky in all its parts. Rhizorne much branched, creeping, rigid and wiry, 2–9 in. long or more; branchlets short, erect, leafy. Leaves tufted, silky on both surfaces, ¼–1 in. long, linear-oblong or linear-obovate, deeply pinnatifid; segments close-set, sometimes slightly recurved, deeply serrate on the upper edge or entire. Peduncles short, stout, rigid, silky, ¼–⅓ in. long. Heads unisexual; males ⅛–⅙ in. diam.; involucral bracts in 1 or 2 series, broadly oblong, silky, with broad purple scarious margins. Florets numerous, eglandular, funnel-shaped. Female heads larger, ⅕–¼ in. diam.; involucral bracts in 3–4 series, larger and broader than in the males, incurved over the florets and often concealing them. Florets numerous; corolla ovoid, inflated at the base, contracted at the minutely toothed mouth. Achenes somewhat curved, rounded on the back or obscurely trigonous.— Kirk, Students' Fl. 327. Leptinella pusilla, Hook. f. Fl. Nov. Zel. i. 129.

North Island: East Cape and Hawke's Bay, Colenso! South Island: Not uncommon throughout. Sea-level to 4500 ft. November–February.

18.C. squalida, Hook. f. Handb. N.Z. Fl. 143.—Stems long, slender, creeping, branched, 4–12 in. long or more, silky or pilose; branchlets short, erect. Leaves 1–2 in. long, linear-obovate in outline, petiolate, flaccid and membranous, more or less pilose with long soft hairs, deeply pinnatifid; segments rather lax, recurved, deeply incised along the upper margin or more rarely along both margins. Peduncles 1–3 in. long, slender, naked, silky. Heads unisexual; males ⅙–⅕ in. diam.; involucral bracts few, in 2 series, broadly oblong-obovate, silky, with broad purplish scarious margins. Florets numerous, eglandular, funnel-shaped. Female heads larger, ¼–⅓ in. diam.; involucral bracts in 3–4 series, broad, rounded, silky, purplish, usually incurved over the florets and concealing them. Florets numerous; corolla ovoid, inflated at the base, much contracted at the mouth, minutely 4-toothed. Achene curved, rounded at the back, almost trigonous.—Kirk, Students' Fl. 328. Leptinella squalida, Hook. f. Fl. Nov. Zel. i. 129.

North Island: Bay of Plenty, Petrie! East Cape and Hawke's Bay, Colenso! Mount Hikurangi, Adams and Petrie! Mount Egmont ranges, T. F. C. South Island: Not uncommon from Nelson to Foveaux Strait. Sea-level to 4000 ft. December–February.

Closely allied to C perpusilla, but distinguished by the greater size, soft flaccid leaves with deeply incised segments, and by the larger pistillate heads, which have the involucral bracts so much incurved as to completely hide the florets.

19.C. dioica, Hook. f. Handb. N.Z. Fl. 143.—Stems creeping, rather stout or slender, glabrous or slightly hairy, 3–12 in. long. Leaves tufted or solitary, membranous, often flaccid, petiolate, page 359½–2 in. long, linear-obovate or spathulate, obtuse, crenate-serrate or lobulate or semipinnatifid, often deeply pinnatifid or pinnate at the base, lobes or segments entire or the upper margins more or less deeply toothed or incised. Peduncles axillary, longer or shorter than the leaves, naked, pubescent. Heads unisexual; males ⅙–¼ in. diam.; involucral bracts few, in 2 series, oblong - orbicular, pubescent, with broad purple scarious margins. Florets numerous, eglandular, funnel-shaped. Female heads larger, ⅕–⅓ in. diam. or more; involucral bracts more numerous, imbricated in 3 or 4 series, incurved. Florets many; corolla ovoid-conic, inflated at the base, narrowed and minutely toothed at the mouth. Achene obovoid, curved, rounded at the back.—Kirk, Students' Fl. 328. Leptinella dioica, Hook. f. Fl. Nov. Zel. i. 129.

North and Souts Islands, Stewart Island: Not uncommon from the Great Barrier Island and the Kaipara Harbour southwards; most plentiful near the sea, but ascending to 3000 ft. November–February.

A most puzzling plant, exceedingly variable in the size of the leaves and the extent to which they are toothed or divided, and also varying much in the size of the flower-heads. Mr. Kirk divided it into three species; but these are clearly connected by intermediate forms, and cannot always be distinguished by the descriptions he has given. The following are the chief varieties, but it must be borne in mind that the characters used to separate them are purely arbitrary.

Var. a.—Leaves 1–2 in., obovate-spathulate, membranous, flaccid, toothed or lobulate above, pinnatifid below, segments entire or toothed. Abundant.

Var. crenatifolia, Kirk, Students' Fl.—Leaves ¾–3 in., oblong-spathulate, membranous, crenate-toothed, with a few deeper divisions at the base of the leaf. Mountain swamps in the South Island.

Var. rotundata.—More robust, hairy. Leaves ½–1½ in., orbicular-spathulate, toothed above, usually lobulate or pinnatifid below.—Cliffs between the Manukau and Kaipara Harbours, T. F. C.; East Coast, Colenso!

Var. pulchella.—Slender. Leaves ⅓–2½ in., linear-oblong or linear-obovate, membranous but firm, pinnatifid, often pinnate at the base. Often difficult to separate from var. a, but. leaves usually narrower and more deeply divided.— C. pulchella, Kirk, Students' Fl. 328. Swamps in the South Island, Stewart Island.

Var. obscura. — Very small, 4/3–1 in. long. Leaves few, ⅓–½ in., oblong-lanceolate or linear-oblong, lobed or pinnatifid, lobes 3 or 4 on each side. Heads small, 1/10–⅛ in. diam.—C. obscura, Kirk, l.c. 327. Swamps at Woodend, Southland, Kirk!