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Manual of the New Zealand Flora.

9. Gnaphalium, Linn

9. Gnaphalium, Linn.

Hoary or woolly annual or perennial herbs. Leaves alternate, quite entire. Heads rather small, corymbose or fascicled or solitary, heterogamous and discoid. Involucre ovoid or campanulate; bracts imbricate in several series, more or less scarious, the inner sometimes with white spreading tips. Receptacle naked or pitted. Florets of the circumference all female, in 2 or more rows, numerous, filiform, minutely 3–4-toothed. Disc - florets hermaphrodite, fewer in number, tubular with a funnel-shaped 5-toothed mouth. Anthers sagittate at the base, produced into fine tails. Style-branches of the disc-florets nearly terete, truncate or capitate. Achenes oblong or obovoid, not ribbed. Pappus-hairs in one series, slender or thickened at the tip, caducous, often connate at the base.

A large genus, spread over the whole world, hardly distinguishable from Helichrysum and others except by the more numerous female flowers. Of the New Zealand species two are widely distributed in many countries, two others extend to Australia, the remainder are endemic.

A. Inner involucral bracts white and radiating. Heads corymbose.

Stems robust. Leaves 2–4 in. x ¼–¾ in., oblong-lanceolate, 3-nerved beneath. Heads ½ in. diam. 1. G. Lyallii.
Stems slender, prostrate or decumbent. Leaves ½–1 in., obovate - spathulate, faintly 3-nerved above. Heads ½–⅓ in. diam. 2. G. trinerve.
Stems slender. Leaves ½–2½ in. x ⅛–½ in., linear-lanceolate. Heads ⅓ in. diam. 3. G. Keriense.
Stems branched, slender, rigid. Leaves ½–1¼ in. x 1/40–1/20 in., narrow-linear, margins recurved. Heads ⅓ in. 4. G. subrigidum.

B. lunar involucral bracts not white, and radiating.

* Heads solitary, terminal.

Small, 1–4 in. high, simple or branched below. Leaves ⅓–2 in., linear-obovate, white on both surfaces. Head ⅓ in. diam. 5. G. Traversii.
Small, 1–2 in. high, solitary or tufted. Leaves ⅓–1 in., linear-spathulate, white beneath. Heads ⅕ in. diam 6. G. paludosum.
Small, stems short, tufted. Leaves imbricate, ⅓ in., linear, silky on both surfaces. Head ½ in. diam. 7. G. nitidulum.

** Heads numerous, corymbose, ebracteate.

Stems 3–18 in. Leaves 1–3 in., linear or linear-spathulate, white and woolly on both surfaces 8. G. luteo-album.

*** Heads compacted into a dense glomerule, with linear bracts at the base.

Annual, stems 6–20 in., branched, erect, leafy. Glomerules ½–1 in. diam. 9. G. japonicum.
Perennial, with creeping stolons. Stems 3–12 in., simple. Leaves mostly radical. Glomerules smaller: 10. G. collinum.
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1.

G. Lyallii, Hook. f.-Fl. Nov. Zel. i. 137.—Stems 1–2 ft. high, stout, branched, decumbent or prostrate at the woody base, then erect, cottony above. Leaves close - set or the upper remote, spreading, 2–4 in. long, ¼–¾ in. broad, narrow oblong-lanceolate or narrow obovate-lanceolate, acute, gradually narrowed to a sessile base, membranous, glabrous above, beneath clothed with thinly appressed white tomentum and distinctly 3-nerved. Heads rather large, in. diam., disposed in many - flowered terminal corymbs 2–4 in. across; pedicels cottony. Involucral bracts numerous, scarious, white, radiating, with short woolly claws. Female florets very numerous, disc-florets much fewer. Achene linear-oblong, smooth, glabrous. Pappus-hairs few, very slender.—Handb. N.Z. Fl. 152; Kirk, Students' Fl. 297. G. adhærens, Col. in Trans. N.Z. Inst. xvii. (1885) 244.

North Island: Ruahine Range, Colenso! Andrew, Howlett! Murimotu and Moawhango, Petrie! Rimutaka Range, Kirk! South Island: Not uncommon along the western side from Collingwood to the Otira Gorge and southwards to Milford Sound. Sea-level to 2500 ft. November–January.

A handsome plant, in its usual state at once recognised by the large size, broad distinctly 3-nerved leaves, and large heads. But small states with narrower leaves and smaller heads are difficult to separate from G. Keriense, and there is little doubt that the two species pass into one another. In both G. Lyallii and Keriense the disc-florets are frequently sterile, showing a tendency in the heads to become unisexual.

2.

G. trinerve, Forst. Prodr. n. 289. —Stems 1–2 ft. long, rather slender, sparingly branched, prostrate or decumbent, glabrate below, more or less white and cottony above. Leaves distant, spreading or deflexed, ½–1 in. long, obovate-lanceolate or spathulate, broadest towards the tip, acute or apiculate, gradually narrowed to a broad base, glabrous above and faintly 3-nerved, white with appressed wool beneath. Heads in corymbs of 3 to 10 or solitary, terminating the branches, which are produced into slender cottony bracteate peduncles, ½–¾ in. diam. Involucral bracts numerous, white, radiating, the outer woolly at the base. Achene linear-oblong, glabrous. Pappus-hairs few, slender.—A. Rich. Fl. Nouv. Zel. 239; A. Cunn. Precur. n. 455; Raoul, Choix, 45; Hook. f. Fl. Nov. Zel. i. 138: Handb. N.Z. Fl. i. 153; Kirk, Students' Fl. 297.

North Island: Rimutaka Range, Kirk! South Island: West coast of Nelson to Westland and Milford Sound, south and east coasts of Otago; not uncommon. Sea-level to 2000 ft. November–December.

Closely allied to the preceding, but sufficiently characterized by the more slender stems, shorter obovate-spathulate leaves, which are seldom 3-nerved beneath, and by the bracteate peduncles.

3.G. Keriense, A. Cunn. Precur. n. 454. — Stems short or long, prostrate or decumbent, with numerous erect or ascending leafy branches 3–9 in. long, cottony above. Leaves spreading, page 324½–1½ in. long, ⅛–½ in. broad, narrow linear-lanceolate to oblong-lanceolate or spathulate-laneeolate, acute, gradually narrowed to the base, glabrous above, beneath white with appressed cottony tomentum but with the midrib evident, rarely faintly 3-nerved. Heads small, ⅓ in. diam., in many-flowered corymbs terminating the branches, which are produced into leafy or bracteate cottony peduncles. Involucral bracts numerous, white, radiating; outer cottony at the base. Achene minute, glabrous. Pappus-hairs few, very slender.—Raoul, Choix, 45; Hook. f. Fl. Nov. Zel. i. 138; Handb. N.Z. Fl. 153; Kirk, Students Fl. 298. G. novæ-zealandiæ, Sch. Bip. in Bot. Zeit. iii. (1845) 171. Helichrysum micranthum, A. Cunn. ex D.C. Prodr. vi. 189.

North Island: Abundant by the sides of streams, &c, from Mongonui to Cook Strait. South Island: Nelson, Travers; Wangapeka Valley, T. F. C.; Dusky Bay, Lyall. Sea-level to 2000 ft. October–December.

4.

G. subrigidum, Col. in Trans. N.Z. hist. xvii. (1885) 245.— Stems usually much branched, sometimes almost bushy, erect or decumbent at the base, woody, 9–20 in. long; branches slender, brittle, somewhat rigid, glabrous or cottony at the tips. Leaves close-set, spreading, ½–1¼ in. long, 1/40–1/15 in. broad, very narrow-linear or narrow linear-spathulate, acute or apiculate, stiff and coriaceous, glabrous above, white with appressed tomentum beneath but with the midrib evident; margins recurved. Heads very numerous, ⅓–½ in. diam., in broad many-flowered corymbs at the ends of the branches; pedicels usually very slender, almost capillary, white and cottony. Involucral bracts white and radiating. Achene and pappus as in G. Keriense.—G. Keriense var. linifolia, Hook. f. Fl. Nov. Zel. i. 138; Handb. N.Z. Fl. 153; Kirk, Students' Fl. 298.

North Island: Dry rocky banks from the East Cape and Taupo to Wauganui; not uncommon. Sea-level to 2000 ft. October–December.

This has generally been treated as a variety of G. Keriense, from which it differs in the more rigid erect and bushy habit, much narrower stiff and coriaceous leaves with recurved margins, and almost capillary pedicels. It deviates quite as much from G. Keriense as Lyallii and trinerve do, and for the sake of consistency should be considered as distinct, unless the four species are merged into one.

5.

G. Traversii, Hook. f. Handb. N.Z. Fl. 154.—A dwarf tufted perennial herb 1–4 in. high, simple or branched below, sometimes forming small patches. Leaves radical, petiolate, ⅓–2; in. long, linear-spathulate or linear-obovate, clothed on both surfaces with soft cottony wool, petiole long or short. Scapes erect, variable in length, ½–3; in., white with cottony wool; bracts 1–3, linear. Head solitary, terminal, ⅓ in. diam.; involucral bracts in about 3 series, erect, linear, obtuse, pale brownish-yellow, scarious and hyaline, shining; outer shorter and broader, cottony at the base. Female page 325florets very numerous, with a few hermaphrodite ones in the centre. Achene puberulous or glabrous. Pappus-hairs copious, very fine, connate at the base.— Benth. Fl. Austral. iii. 655; Kirk, Students' Fl. 299.

Var. Mackayi, T. Kirk, l.c.—Much branched, densely tufted, forming close matted patches. Leaves imbricated on the short branches, heads smaller, sessile or on short scapes. Involucral bracts not so numerous, acute.—Raoulia Mackayi, Buch, in Trans. N.Z. Inst. xiv. (1882) 354, t. 34, f. 2.

South Island: Not uncommon in mountain districts from Nelson to Foveaux Strait. 1500–5500 ft. December–February. Also in Australia.

A variable plant. The most developed form is almost simple, with long radical leaves and long and slender scapes; but it passes by almost imperceptible gradations into the extreme state of var. Mackayi, which forms broad much-branched patches, with short imbricated leaves and sessile heads.

6.G. paludosum, Petrie in Trans. N.Z. Inst. xxii. (1890) 441. —Very slender, solitary or tufted, 1–2 in. high. Leaves all radical, petiolate, ⅓–1½ in. long, narrow linear-spathulate; blade half the length or nearly so, acute or subacute, rather membranous, glabrous or slightly silky above, beneath clothed with white appressed tomentum; midrib prominent; margins flat or slightly recurved. Scapes few, hardly exceeding the leaves in the flowering stage, but elongating in fruit, very slender, almost capillary, cottony; bracts few, small, linear. Head solitary, terminal, ⅕ in. diam.; involucral bracts few, scarious, pale and glistening, darker at the tips, inner linear, glabrous. Female florets very numerous. Aehene linear-oblong, papillose. Pappus-hairs very delicate, connate at the base. —Kirk, Students' Fl. 299. G. minutulum, Col.in Trans. N.Z. Inst. xxii. (1890) 472.

North Island: Base of Tongariro, Hill! Rangipo Plains and Ruahine Mountains, Petrie! South Island: Not uncommon in mountain districts, usually in peaty swamps, from Nelson to Otago. 1000–4000 ft. December–January.

Closely allied to G. Traversii, but easily separated by the smaller size and more slender habit, thinner leaves glabrous on the upper surface, smaller heads, and fewer involucral bracts with dark tips. The type specimens of Mr. Golenso's G. minutulum are identical with Mr. Petrie's plant.

7.G. nitidulum, Hook. f. Handb. N.Z. Fl. 154.—" A small densely tufted species, covered with appressed silky shining yellowish tomentum. Leaves closely imbricated at their bases, above spreading, flat, ⅓ in. long, linear, obtuse; lower ⅓ membranous, glabrous, upper ⅔ densely silky. Heads terminal, solitary, large, ½ in. broad, on very short slender peduncles; involucral scales in 2 series, erect, linear, hyaline, shining, with pale erect tips; florets not seen."—Kirk, Students' Fl. 299.

South Island: "Nelson Mountains, Sinclair; Clarence and Wairau Valleys, alt. 3500 ft., Travers."

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This plant has apparently not been collected since its original discovery nearly fifty years ago. Not having seen specimens, I have reproduced Hooker's description.

8. G. luteo-album, Linn, Sp. Plant. 851.—Annual or rarely biennial, clothed in all its parts with soft white woolly tomentum. Stems erect, ascending or decumbent, simple or branched at the base, 3–18 in. high. Lower leaves often petiolate, 1–3 in. long, linear or linear-spathulate to obovate-spathulate, obtuse or acute; upper smaller, sessile, linear or lanceolate, acute. Heads ⅕ in. diam., pale-yellow or brownish-yellow, glistening, in dense ebrac-teate clusters arranged in a corymbose manner at the ends of the branches. Involucre almost globose; bracts erect, oblong, obtuse, scarious, tips incurved. Female florets exceedingly numerous; hermaphrodite ones few. Achene minutely papillose.—A. Rich. Fl. Nouv. Zel. 236; A. Cunn. Precur. n. 450; Raoul, Choix, 45; Hook. f. Fl. Nov. Zel. 138; Handb. N.Z. FL 154; Kirk, Students' Fl. 298.

Keramadec Islands, North and South Islands, Stewart Island, Chatham Islands, Auckland Islands: Abundant throughout from sea-level to fully 3000 ft. November–March.

A common plant in almost all warm and many temperate countries.

9.G. japonicum, Thunb. Fl. Jap. 311.— Annual, erect, 6–20 in. high. Stems often woody at the base, branched, more or less white and cottony. Leaves scattered, 1–4 in. long, oblong-spathulate or linear-spathulate to linear-lanceolate, the lower often petiolate, acute, glabrous above or nearly so, cottony-white beneath. Heads small, ⅙–¼ in. long, compacted into dense globose clusters or compound heads ¼–1 in. diam., which either terminate the branches or are axillary, and are surrounded by 3–6 linear spreading floral leaves. Involucral bracts scarious, hyaline, erect, obtuse or the inner acute. Female florets very numerous; hermaphrodite ones few, sometimes solitary. Achene slightly compressed, glabrous. Pappus-hairs very slender, hardly connate at the base.—Benth. Fl. Austral. iii. 653; Kirk, Students' Fl. 300. G. involucratum, Forst. Prodr. n. 291; A. Rich. Fl. Nouv. Zel. 241; A. Cunn. Precur. n. 453; Raoul, Choix, 45; Hook. f. Fl. Nov. Zel. i. 139; Handb. N.Z. Fl. 155. G. virgatum, Banks and Sol. ex Hook. f. Fl. Nov. Zel. i. 139. G. lanatum, Forst. Prodr. n. 290; A. Cunn. Precur. n. 452. G. Cunninghamii, D.C. Prodr. vi. 235.

Kermadec Islands, North and South Islands, Stewart Island, Chatham Islands: Abundant throughout from sea-level to 2500 ft. November–January.

Found also in Norfolk Island, Lord Howe's Island, Australia and Tasmania) and northwards to the Malay Archipelago, China, and Japan.

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10.G. collinum, Lab. PL Nov. Holl. ii. 44, t 189.—A tufted perennial herb 3–12 in. high, usually with creeping and rooting stolons and slender erect cottony stems. Leaves mostly radical, very variable in size, ⅓–3 in. long, lanceolate-spathulate or oblong-spathulate, acute or obtuse, petiolate, white and cottony on both surfaces or glabrate above; cauline leaves much smaller and narrower, linear - spathulate, sessile. Heads small, compacted into dense clusters or compound heads similar to those of G. japonicum, but smaller and not so compact, and with fewer smaller subtending floral leaves. Involucres broader than in G. japonicum; bracts linear-oblong, obtuse, scarious and hyaline. Florets and achenes as in G. japonicum.—Hook. f. Fl. Nov. Zel. i. 139; Handb. N.Z. FL 155; Benth. Fl. Austral. ii. 654; Kirk, Students' Fl. 300. G. simplex, A. Rich. Fl. Nouv. Zel. 237; A. Cunn. Precur. n. 451.

Var. obscurum. Kirk, Students' Fl. 300.—Forming grey matted patches seldom more than 1 in. high. Leaves ½–¾ in., linear, glabrate above, grey with appressed tomentum beneath. Scape leafy or almost wanting. Heads in terminal fascicles of 2–4 or solitary. Involucral bracts 3–10, linear, obtuse.

Var. monocephalum, Kirk, l.c.—Very small. Leaves all radical, ⅓ in. long, linear, obtuse or acute. Head solitary, sessile or on a slender filiform scape ½–1 in. high.

Kermadec Islands, North and South Islands, Stewart Island, Chatham Islands: Plentiful from sea-level to 4500 ft. November–March. The two varieties not uncommon on the mountains of the South Island.

Luxuriant forms of this are best distinguished from G. japonicum by being perennial, by the creeping stolons, smaller glomerules with fewer floral leaves, and by the broader involucres. It is also a native of Australia and Tasmania.