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Manual of the New Zealand Flora.

16. Lomaria, Willd

16. Lomaria, Willd.

Rhizome creeping or short and suberect, sometimes lengthened into a short caudex, rarely long and climbing. Fronds variable in size, usually simply pinnate or pinnatifid, rarely undivided, very rarely bipinnate, dimorphic; the outer fronds sterile with large and broad flat pinnæ the inner fertile with smaller linear pinnæ. Veins free, not anastomosing. Sori linear, in a continuous elongated line occupying the whole space between the midrib and the margin. Indusium linear, membranous, composed of the more or less modified edge of the frond, at first revolute over the sorus, ultimately spreading. Sporangia stalked, girt by an incomplete vertical ring, bursting transversely.

A large genus of nearly 50 species, most abundant in the south temperate zone, but with outlying species in most temperate and tropical countries. It only differs from Blechnum in the sori being close to the margin of the frond, and is united with that genus by many pteridologists. Of the 14 speoies found in New Zealand 4 or perhaps 5 are endemic, 2 are widely distributed in the south temperate zone, the remainder are found either in Australia or the Pacific islands, or in both.

A. Fronds pinnate or pinnatifid, rarely simple.

* Sterile fronds pinnatifid (or rarely simple), central and lower pinna connected by their dilated bases.

Fronds 1–3 ft. long, often pendulous, broad, coriaceous, sometimes simple. Pinnæ few, large, 4–12 in. long, 1–1 ½ in. broad 1. L. Patersoni.
Fronds 1–4 ft. high, narrow, erect. Pinnæ very numerous, 1–3 in. X ¼–½ in., reddish or dirty-white beneath 2. L. discolor.
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** Sterile fronds pinnatifid above, pinnate below, central and lower pinnæ free, but mere or less dilated at their bases.

a. Pinnæ at the base of the frond not reduced in size. Fronds 4–14 in., lanceolate-deltoid. Pinnæ 1–3 in., lanceolate or ensiform, the lowest pair often deflexed 3. L. vulcanica.

b. Pinnæ gradually reduced in size towards the base of the frond.

Fronds 1–3 ft. × 3–6 in., submembranous. Pinnæ l ½–3 in. × ⅓–⅔ in., lanceolate, falcate, acuminate. Pinnæ of fertile fronds 2–3 in. long 4. L. Norfolkiana.
Fronds 6–18 in. × 2–4 in., submembranous. Pinnæ 1–2 in, × ¼–½ in., oblong-lanceolate, obtuse or acute, sinuate-crenate. Pinnæ of fertile fronds ¾–1½ in., narrow-linear, acute 5. L. lanceolata.
Fronds 9–30 in. × 1½–4 in., fleshy or coriaceous. Pinnæ ¾–2½ in. × ¼–½ in., linear-oblong to lanceolate, entire. Pinnæ of fertile fronds ¾–1½ in., linear-oblong, obtuse 6. L. dura.
Rhizome short. Fronds 4–12 in. × ½-l in. dark-green, coriaceous. Pinnæ ⅓–½ in. × ¼ in., oblong to suborbicular. Fertile fronds shorter than the sterile 7. L. L Banksii.
Rhizome creeping. Fronds 2–12 in. × ⅓–⅔ in., coriaceous to submembranous. Pinnæ ⅕–⅓ in., ovate-oblong to linear-oblong. Fertile fronds longer than the sterile; pinnse linear-oblong, obtuse 8. L. alpina.

*** Sterile fronds pinnate, or pinnatifid above; Pinnæ not dilated at their

Rhizome short, thick. Fronds often very large, 1–8 ft. long. Pinnæ 3–12 in. × ½-l in., linear, coriaceous 9. L. capensis.
Rhizome very long, climbing. Sterile fronds dimorphic: lower with small rounded Pinnæ; upper with long lanceolate falcate ones 10. L. filiformis.
Rhizome short, suberect. Fronds 3–8 in. × 1–1½ in., blackish-green, lyrate-pinnatifid. Terminal Pinnæ much longer than the lateral 11. L. nigra.
Rhizome stout, suberect. Fronds 12–30 in. × ¾–1½ in., linear, membranous; rhachis and stipes bristly and scaly. Pinnæ ½–¾ in. × ¼–⅓ in., oblong to suborbicular, obtuse 12. L. fluviatilis.
Rhizome short, suberect. Fronds 3–10 in. × ¾–1½ in., submembranous; stipes and rhachis naked. Pinnæ ½–¾ in., ovate-oblong to oblong, obtuse 13. L.membranacea.

B. Fronds bipinnate.

Rhizome often produced into a caudex resembling the trunk of a miniature tree-fern. Fronds 9–18 in. long, ovate, acuminate 14. L. Fraseri.
1.L. Patersoni, Spreng. Syst. Veg. iv. 62; var. elongata, Hook. and Bah. Syn. Fil. 174.—Rhizome short, stout, creeping, clothed with blackish-brown scales, sometimes stoloniferous; rootlets tomentose. Stipes 3–9 in. long, stout, black, scaly at the base. Sterile fronds very variable; of young plants (and occasionally of old ones) quite simple and entire, 6–12 in. long, 1–1½ in. broad; of old plants pinnatifid, 1–3 ft. long or more, 6–12 in. broad, broadly page 976ovate to ovate-lanceolate, erect or pendulous, coriaceous, glabrous, dark-green above, paler beneath; rhachis winged throughout. Pinnæ 4–12 on each side, alternate, acuminate, broadly decurrent at the base, forming a rounded lobe in the sinus, quite entire; at the base of the frond there are usually several small rudimentary Pinnæ sometimes extending down the stipes almost to its base. Veins numerous, close, free, forked. Fertile fronds as long as the sterile, pinnatifid; Pinnæ narrow-linear, 4–10 in. long, ⅛–¼ in. broad. Sori continuous, ultimately covering the whole under-surface except the costa.—Thoms. N.Z. Ferns, 64; Field, N.Z. Ferns, 98, t. 11, f. 3, 3A, 3B. L. elongata, Blume, En. Fil. Jav. ii. 201; Hook. Sp. Fil. iv. 3, t. 143; Hook. f. Fl. Nov. Zel. ii. 29; Handb. N.Z. Fl. 367. L. heterophylla, Col. in Tasmanian Journ. Nat. Sci. (1845) 15 (not of Desv.). L. Colensoi, Hook. f. Ic. Plant, t. 627, 628. Blechnum Patersoni, Metten.

North and South Islands, Stewart Island: Damp hilly forests from the Thames and Te Aroha southwards, not common; local on the east side of the South Island. Sea-level to 3000 ft.

The New Zealand variety is also found in the Pacific islands, Malaya, and India. The typical state, which differs in the fronds being usually simple, occurs in Australia, Tasmania, and the Philippines. As in most of the species of the genus, the fronds are sometimes partly fertile and partly sterile.

2.L. discolor, Willd. Sp. Plant. v. 293.—Rhizome short, stout, suberect, stoloniferous at the base, often lengthened above into a short erect caudex 1–2 ft. high, clothed at the top with the bases of the old stipites. Stipes 3–6 in. long, stout, polished, densely covered at the base with dark-brown linear scales. Fronds numerous, tufted at the top of the caudex and forming an elegant crown, erect, 1–4 ft. high; sterile linear-lanceolate to oblong-lanceolate, gradually tapering at both ends, 2–6 in. broad in the middle, coriaceous, glossy-green above, dirty-white to reddish-brown beneath, often clothed with rufous scales when young, glabrous when old, deeply pinnatifid or pinnate at the very base. Pinnæ very numerous, closely placed, horizontally spreading, 1–3 in. long, ¼–½ in. broad, linear to linear-oblong, subacute, usually connected by their broad dilated bases, margins minutely sinuate. Veins close, free, forked. Fertile fronds about as long as the sterile but narrower; Pinnæ ¾–1½ in. long, linear, stout, often flexuous, usually with broad leafy bases. Sori continuous, covering the whole under-surface except the costa. Indusium with the margins much lacerated.—A. Cunn. Precur. n. 181; Raoul, Choix, 37; Hook. Sp. Fil. iii. 5; Hook. f. Fl. Nov. Zel. ii. 30; Handb. N.Z. Fl 368; Hook, and Bak. Syn. Fil. 175; Benth. Fl. Austral. vii. 735; Thoms. N.Z. Ferns, 65; Field, N.Z. Ferns, 100, t. 4, f. 2, 2A. Stegania discolor, A. Rich. Fl. Nouv. Zel. 87. Onoclea discolor, Swartz, Syn. Fil. iii. Osmunda discolor, Forst. Prodr. n. 413. Blechnum discolor, Metten.page 977

North and South Islands, Chatham Islands, Stewart Island, Auckland and Campbell Islands: Abundant in open forests throughout. Sea-level to 3000 ft.

Easily distinguished by the tall erect habit, long and narrow horizontally spreading pinnæ, and dirty-white or reddish under-surface. The fronds are frequently forked at the top, and a beautiful sport is in cultivation in which the pinnæ are greatly expanded in the upper two-thirds of their length, and deeply pinnatifid. Also a native of Norfolk Island, Australia, and Tasmania.

3.L. vulcanica, Blume, En. Fil. Jav. ii. 202.—Rhizome short, stout, woody, erect or inclined, densely clothed with the remains of the old stipites. Stipes 4–9 in. long, slender, pale yellow-brown, clothed towards the base with dark-brown shining subulate scales, smooth and polished above. Sterile fronds 4–14 in. long without the stipes, 2–5 in. broad at the base, lanceolate-deltoid, not narrowed below, acuminate, coriaceous, dull-green, glabrous or the surfaces and margins sprinkled with soft white hairs, pinnate at the base, pinnatifid above. Pinnæ 1–3 in. long, ¼–⅓ in. broad, spreading, lanceolate or ensiform, broadest at the base, acute or obtuse at the tip, falcate, lowest pair deflexed; margins thickened, entire or minutely crenate-undulate. Veins free, forked. Fertile fronds usually exceeding the sterile and with a longer stipes, pinnate in the lower half; Pinnæ 1–2 in. long, linear, distant, with a dilated adnate base. Sori continuous; indusium with lacerate margins.— Hook. Ic. Plant, t. 969; Sp. Fil. iii. 12; Hook. f. Fl. Nov. Zel. ii. 29; Handb. N.Z. Fl. 367; Hook. and Bak. Syn. Fil. 176; Benth. Fl. Austral. vii. 735; Thoms. N.Z. Ferns, 65; Field, N.Z. Ferns, 99, t. 27, f. 5, 5A. L. deitoides, Col. in Tasmanian Journ. Nat. Sci. (1845) 17. L. deflexa, Col. l.c. 18. L. paucijuga, Col. in Trans. N.Z. Inst. xx. (1888) 222. Blechnum vulcanicum, Christ.

North and South Islands, Stewart Island: In dry open woods from Auckland and Coromandel southwards, but often rare and local, especially to the north of the East Cape, more frequent in the subalpine forests of Nelson and Canterbury. Sea-level to 3500 ft.

A well-marked species, at once recognised by the narrow-deltoid frond, with the lowest pair of pinnse deflexed. It extends northwards through Australia and the Pacific islands to Malaya.

4.L. Norfolkiana, Heward in Lond. Journ. Bot. (1842) 122.—Rhizome short, stout, erect or inclined, clothed with the bases of the old stipites mixed with dark-brown chaffy scales. Stipes short, stout, 2–4 in. long, scaly at the base. Sterile fronds numerous,. forming a crown at the top of the rhizome, erect or spreading, 1–3 ft. high, 3–6 in. broad, lanceolate or narrow elliptic-lanceolate, gradually tapering from the middle to both ends, acuminate, dark-green, firm but scarcely coriaceous, quite glabrous, deeply pinnatifid or pinnate at the base. Pinnæ numerous, close-set, horizontally spreading, 1½–3 in. long, ⅓–⅔ in. broad, lanceolate, tapering from a broad adnate base to an acuminate point, subfalcate, the lower ones page 978gradually reduced in size to minute auricles, margins crenulate; veins fine, close, forked. Fertile fronds rather shorter than the sterile, pinnate; pinnæ remote, very narrow-linear, 2–3 in. long, apiculate.—Bak. in Ann. Bot. v. (1891) 219. L. acuminata, Bak. Syn. Fil. (edit. 2) 481; Thoms. N.Z. Ferns, 66. L. attenuata, Hook. f. Handb. N.Z. Fl. 368; Field, N.Z. Ferns, 101 (not of Willd.).

Kermadec Islands: Sunday Island, abundant, McGillivray, T. F. C. North Island: Three Kings Islands, T. F. C. Little Barrier Island, Reischek! T. F. C.

Also in Norfolk Island. It can only be distinguished from L. lanceolata by the greater size, the long acuminate sterile pinnæ, and the much longer fertile pinnæ, and might well be regarded as a variety only. On both the Three Kings Islands and the Little Barrier it appears to gradually merge into the ordinary state of L. lanceolata.

5.L. lanceolata, Spreng. Syst. Veg. iv. 62.—Bhizome short,. stout, erect or inclined, rarely produced into a short caudex 3–6 in. high. Stipes 2–6 in. long, firm, erect, dark-brown at the base and clothed with subulate scales, paler and glabrous above. Fronds. tufted, forming a crown at the top of the rhizome; the sterile ones 6–18 in. long, 2–4 in. broad, lanceolate, acuminate, gradually narrowed to the base, rather membranous, pale-green or dark-green, quite glabrous, pinnate below, pinnatifid above. Pinnæ numerous, close-set, horizontally spreading or ascending, 1–2 in. long, ¼–½ in. broad, lanceolate or oblong-lanceolate, attached by a broad somewhat dilated base, gradually tapering to an obtuse or acute point, slightly falcate, usually sinuate-crenate towards the tip, rarely entire; veins conspicuous, free, forked. Fertile fronds usually shorter than the sterile, 1–2 in. broad, pinnate; Pinnæ ¾–1 ½ in. long, distant, narrow-linear, acute or apiculate.—A. Cunn. Precur. n 180; Raoul, Choix, 37; Hook. Ic. Plant, t. 429; Sp. Fil. iii. 11; Hook. f. Fl. Nov. Zel. ii. 29; Handb. N.Z. Fl. 367; Hook, and Bak. Syn. Fil. 177; Benth. Fl. Austral. vii. 735; Thoms. N.Z. Ferns, 66; Field, N Z. Ferns, 102, t. 11, f. 2, 2A. L. aggregata, Col. in Trans. N.Z. Inst. xx. (1888) 223; Field,. Z. Ferns, 103, t. 29, f. 7, 7a. Blechnum lanceolatum, Sturm.

North and South Islands, Chatham Islands, Stewart Island: From the North Cape southwards, abundant by the margins of streams, &c. Sea-level to 2500 ft.

Also in Victoria, Tasmania, South Australia, and the Pacific islands. A variable plant. Large specimens pass into L. Norfolkiana, and smaller ones are sometimes difficult to separate from L. membranacea.

6.L. dura, Moore in Gard. Chron. (1866) 290.—Rhizome stout, erect, clothed with the bases of the old stipites, sometimes lengthened into a short caudex. Stipes 1–2 in. long, clothed at the base with large ovate-lanceolate brownish scales. Fronds. page 979numerous, tufted, forming a crown at the top of the rhizome; sterile 1–2 ½ ft. long, 1½–4 in. broad, lanceolate, acute or acuminate, usually broadest above the middle, very gradually narrowed to the base, dark-green, fleshy or almost coriaceous, pinnatifid above, pinnate below. Pinnæ numerous, close-set, often overlapping, the largest ¾–2½ in. long, ¼–½ in. broad, variable in shape, linear-oblong to lanceolate, obtuse or acute, often falcate, attached by a broad base, the upper narrower and more acute, lowermost dwarfed to rounded auricles; margins entire, slightly thickened. Veins free, forked. Fertile fronds shorter and narrower than the sterile, 1–2 ½ in. broad; pinnæ numerous, close, linear-oblong, rigid, obtuse. Sori very copious, covering the whole under-surface.—Hook, f. Handb. N.Z. Fl. 748; Hook, and Bak. Syn. Fil. 177; Thoms. N.Z. Ferns, 66; Field, N.Z. Ferns, 104, t. 10, f. 4, 4A. L. rigida, J. Sm. Ferns, Brit, and For. 290.

South Island: Banks Peninsula, Armstrong. Eastern and southern coasts of Otago, not uncommon, Petrie! Thomson, Kirk! West Coast sounds, J. D. Enys! Stewart Island and The Snares: Kirk! Chatham Islands: Abundant, Chudleigh! Buchanan! Miss Seddon! Auckland and Campbell Islands, Antipodes Island: Kirk!

A purely littoral plant, never found far from the influence of sea-spray. Easily recognised by the fleshy or coriaceous habit, the usually obtuse and entire sterile pinnæ, and by the close-set broad and rigid fertile pinnæ.

7.L. Banksii, Hook. f. Fl. Nov. Zel. ii. 31, t. 76.—Rhizome short, stout, woody, erect or inclined, clothed with the old stipites at the top, and with matted fibres below. Stipes short, stout, dark-coloured, furnished at the base with numerous ovate-lanceo-late chaffy scales. Fronds numerous; sterile 4–12 in. high, rarely more, ½–1 in. broad, linear-lanceolate, narrowed to both ends, rather coriaceous, dark-green, sometimes with a glaucous tinge, pinnatifid above, pinnate below. Pinnæ numerous, close-set, adnate by a broad base, ⅓–½ in. long, rarely more, about ¼ in. broad, broadly oblong or almost semi-orbicular, obtuse, quite entire; lower pinnæ much reduced, sometimes forming a sinuated wing down to the base of the stipes. Fertile fronds usually shorter than the sterile, pinnate throughout; pinnæ shorter and narrower, more distant, straight or curved. Sori copious, covering the whole under-surface.—Handb. N.Z. Fl. 368; Hook. Sp. Fil. iii. 17; Hook, and Bak. Syn. Fil. 178; Thoms. N.Z. Ferns, 67; Field, N.Z. Ferns, 105, t. 26, f. 2, 2a. Blechnum Banksii, Mettenius.

North Island: Auckland—North Cape, Buchanan; near Ahipara, T. F. C.; Bay of Islands, A. Cunningham; Little Barrier Island, Kirk! T. F. C.; Manukau Heads, Sinclair, Colonel Haultain! East Cape, Colenso! Taranaki—White Cliffs to Cape Egmont, Buchanan, T. F. C. Wellington—Wellington Heads, Field; Cape Terawiti, Kirk! South Island: Marl-borough—Queen Charlotte Sound, Banks and Solander. Nelson—Cape Fare-well, Kirk! West Wanganui, Kingsley. Canterbury—Banks Peninsula, page 980Armstrong. Otago—Not uncommon on both the East and West Coasts, Buchanan, Petrie! Thomson, Hamilton! Stewart Island: Paterson's Inlet, Kirk.

A well-marked plant, easily recognised by the coriaceous habit and short and broad rounded Pinnæ attached by a broad base. Like L. dura, it is a purely littoral plant, never found beyond the influence of the sea-spray.

8.L. alpina, Spreng. Syst. Veg. iv. 62.—Rhizome long, slender, branched, creeping, clothed with chaffy ferruginous scales. Stipes 2–6 in. long or more, slender, red-brown, smooth and polished, sparingly scaly. Fronds tufted along the rhizome; sterile shorter than the fertile, 4–18 in. long including the stipes, ⅓–⅔ in. broad, often spreading or decumbent, linear or linear-lanceolate, narrowed to the base, dark-green, pinnatifid or pinnate towards the base, texture varying from thick and coriaceous to almost membranous. Pinnæ numerous, close-set, short, spreading, ⅕–⅓ in. long, attached by a broad base, ovate-oblong or triangular-oblong to linear-oblong, obtuse, entire or obscurely crenate. Fertile fronds erect, pinnate throughout; pinnæ numerous, rather distant, shorter and narrower than the sterile, linear or linear-oblong, obtuse, spreading or deflexed or sometimes curved upwards. Sori copious, covering the-whole under-surface.—Hook. Fil. Exot. t. 32; Sp. Fil. iii. 16; Hook. f. Fl. Antarct. ii. 393, t. 150; Fl. Nov. Zel. ii. 30; Handb. N.Z. Fl. 368; Hook. f. Bak. Syn. Fil. 178; Benth. Fl. Austral. vii. 736; Thoms. N.Z. Ferns, 66; Field, N.Z. Ferns, 105, t. 17, f. 5, 5A. L. pumila, Raoul, Choix, 9, t. 2A; Hook. Sp. Fil. iii. 17; Hook. f. Fl. Nov. Zel. ii. 28; Handb. N.Z. Fl. 367. L. linearis, Col. in Tasmanian Journ. Nat. Sci. (1845) 16. L. parvifolia, Col. in Trans. N.Z. Inst. xx. (1888) 224. Stegania alpina, R. Br. Prodr. 152. Blechnum alpinum, Metten. Fil. Hort. Bot. Lips. 64. Poly-podium penna-marina, Poir. in Lam. Encycl. v. 520.

North and South Islands, Chatham Islands, Stewart Island, Antipcdes Island, Macquarie Island; From the Upper Thames Valley and Rotorua southwards, abundant to the south of the East Cape. Sea-level to 4000 ft.

Also abundant in temperate South America, Australia, and Tasmania. Raoul's L. pumila differs from the type in the more membranous fronds and distinctly crenate pinnæ, but is without doubt a trivial state produced by growing in an unusually sheltered and shaded locality. Specimens exactly resembling Raoul's plate can be found without any difficulty in both islands, and can generally be traced on the spot into ordinary L. alpina. I look upon it as a form too inconstant to keep up even as a variety. L. parvifolia, Col., of which I possess a type specimen forwarded by Mr. Colenso himself, is clearly the same, a view which is also taken by Mr. Baker (Ann. of Bot. v. (1891) 220).

9.L. capensis, Willd. Sp. Plant. v. 291.—Rhizome short, stout, often woody, erect or inclined, sometimes prostrate, clothed at the top with large chestnut-brown scales. Stipes stout, long or short, usually densely scaly at the base. Fronds numerous, very variable in size, usually from 1–4 ft., but in dry exposed places page 981often dwarfed to a few inches, while on the sides of deep wooded ravines they are occasionally 8–10 ft. long or even more; sterile ovate or oblong-ovate to oblong-lanceolate, erect or pendulous, very coriaceous to almost membranous, bright-green to brownish-green, pinnate throughout; rhachis stout, more or less scaly, especially when young. pinnæ often very numerous, but in small specimens and in var. minor frequently reduced to 4–6 pairs, alternate, horizontally spreading, 3–12 in. long or more, ½–1 in. broad, acute or acuminate, oblique at the base and cuneate or truncate or rounded-cordate or even auriculate, sessile by the midrib alone or the uppermost more or less adnate; margins minutely toothed; costgæ more or less scaly. Veins free, close, parallel, usually forked at the base. Fertile Pinnæ very narrow-linear, distant, 3–9 in. long, ⅛–¼ in. broad, usually on separate fronds, but often mixed with sterile pinnse or the Pinnæ partly fertile and partly sterile. Indusium broad, membranous, lacerate.—F. Muell. Veg. Chath. Is. 72; Benth. N.Z. Austral. vii. 737. L. procera, Spreng. Syst. Veg. iv. 65; A. Cunn. Precur. n. 182; Raoul, Choix, 37; Hook. Ic. Plant. t. 427, 428; Sp. Fil. iii. 22; Garden Ferns, t. 53; Hook. f. Fl. Antarct. i. 110; Fl. Nov. Zel. ii. 27; Handb. N.Z. Fl. 366; Hook. and Bak. Syn. Fil. 179; Thoms. N.Z. Ferns, 67; Field, N.Z. Ferns, 107, t. 2, f. 1, 1A. L. latifolia, Col. in Tasmanian Journ. Nat. Sci. (1845) 15. L. duplicata, Potts in Trans. N.Z. Inst. ix. (1877) 491. Stegania procera, R. Br. Prodr. 153; A. Rich. Fl. 86, t. 13. Osmunda capensis, Linn. Mant. 306. O. procera, Forst. Prodr. n. 414. Blechnum capense, Schlecht. Adumb. Fil. 34, t. 18.

Var. a, Hook. f. Fl. Nov. Zel. ii. 27.—Usually tall and robust. Sterile pinnsæ truncate or broadly cuneate at the base.

Var. b, Hook. f. l.c.—Usually tall and robust. Sterile pinnsæ cordate or auriculate at the base.

Var. c, Hook. f. l.c.—Usually tall and robust. Sterile pinnæ narrowed at the base.

Var. d, minor, Hook. f. l.c.—Smaller, 1–3 ft. high, dark olive-green; fertile fronds usually exceeding the sterile. Sterile pinnæ few, 4–8 pairs, short, broad, linear-oblong, the lowermost hardly shorter than the one above it, upper often adnate.—Stegania minor, E. Br. Prodr. 153.

Kermadec Islands, North and South Islands, Chatham Islands, Stewart Island, Auckland and Campbell Islands, Antipodes Island: Abundant throughout, ascending to 4000 ft.

A very widely distributed species. From Australia and Tasmania it extends northwards to Malaya, and is common in many of the Pacific islands. In America it ranges from the south of Chili northwards to Mexico and the West Indies. It is also found in South Africa. In New Zealand it occurs in all soils and situations, and, although attaining its greatest luxuriance in deep forest ravines, is plentiful in open swamps and gullies, and even not averse to bare hillsides or the clefts of rocky peaks. At first it is difficult to believe that the small forms found in exposed places, often not more than 6 in. high, with 3–4 pairs of pinnæ, can belong to the same species as the huge specimens growing on moist cliffs in shaded ravines, in which the fronds are sometimes 8–10 ft. long, with more than 40 pairs of pinnæ. But every gradation of size exists, page 982and one form can be traced directly into the other. I have kept up the four varieties established by Sir J. D. Hooker in the Flora, although the first three do not seem to be separated by any well-defined characters. Var. minor is more distinct; and in some respects approaches L. vulcanica. It has a different habit and mode of growth, and may prove to be a separate species.

In nearly all the species of Lomaria the fertile fronds are sometimes irregularly mixed with sterile pinnæ, but in none is this so commonly seen as in L. capensis. Sometimes one side of the frond may be fertile and the opposite side sterile, or the sterile and fertile pinnæ may be irregularly mixed. Or sometimes the upper half of the frond may be fertile and the lower sterile, or vice versâ. It is also quite common for the pinnæ themselves to be partly fertile and partly sterile. The frond is also occasionally once or twice dichotomously forked, constituting Mr. Potts's L. duplicata, and sometimes the tips of the fronds are regularly crested.

10.L. filiformis, A. Cunn. Precur. n. 183.—Rhizome long, stout, branched, climbing up trees to a great height, clothed with squarrose scales. Sterile fronds very numerous, scattered along the rhizome, pinnate throughout, of two forms; those on the ground or on the lower part of the rhizome small, 3–6 in. long, ½–1 in. broad, linear or linear-lanceolate; pinnæ ¼–½ in. long, oblong to orbicular-oblong, sharply and deeply toothed. Fronds from the upper part of the rhizome much larger, 1–2 ½ ft. long, 3–6 in. broad, lanceolate, pendulous, hardly coriaceous, dark-green, glabrous or more or less scaly along the rhachis and costæ; stipes short, scaly at the base. pinnæ numerous, 1 ½–4 in. long, about ½ in. broad, lanceolate, falcate, narrowed upwards into a finely acuminate point, shortly stipitate and truncate or rounded or cordate at the base, margins regularly and finely crenate-dentate. Fertile fronds from near the top of the rhizome, ovate or ovate-oblong in outline; pinnæ numerous, 3–6 in. long, ⅛ in. broad, very narrow-linear or almost filiform. Indusium very narrow.—Raoul, Choix, 37; Hook. Sp. Fil. iii. 33, t. 149; Hook. f. Handb. N.Z. Fl. 366; Hook, and Bak. Syn. Fil. 180; Thoms. N.Z. Ferns, 68; Field, N.Z. Ferns, 109, t. 10, f. 1 3, 3a, 3b. L. propinqua, A.Cunn. Precur. n. 184. L. pimpi-nellifolia, Hook. f. in Hook. Lond. Journ. Bot. iii. (1844) 412. Stenochlæna heteromorpha, J. Sm. in Hook. Lond. Journ. Bot. iv. (1845) 149; Hook. f. Fl. Nov. Zel. ii. 46; Brack. Fil. U.S. Expl. Exped. 77. Osmunda reptans, Banks and Sol. ex Hook. f. Fl. Nov. Zel. ii. 46. Blechnum reptans, Christ.

North and South Islands: In forests from the North Cape southwards to Nelson and Marlborough, abundant. Sea-level to 2000 ft.

Also recorded from the Fiji Islands. A most distinct species, remarkable for its very long climbing rhizome and dimorphic sterile fronds.

11.L. nigra, Col. in Tasmanran Journ. Nat. Sci. (1845) 16.— Rhizome short, stout, suberect, clothed with the bases of the old stipites mixed with chaffy scales. Stipes slender, densely scaly, 1–3 in. long. Sterile fronds tufted at the top of the rhizome, spreading, 3–8 in. long, 1–1 ½ in. broad, linear-oblong, membranous, page 983blackish-green or lurid-green, brittle when dry, glabrous or the margins and under-surface more or less clothed with short rufous hairs, lyrate-pinnatifid, pinnate at the base; rhachis usually densely pubescent. Pinnæ 4–8 pairs, unequal in size; the terminal one much the largest, 1–2 in. long, oblong, obtuse, irregularly lobed or sinuate; the lateral ¼–¾ in. long, oblong to orbicular-oblong, irregularly sinuate, the lowest pair larger than those immediately above, and often stipitate and deflexed. Fertile fronds few, erect, pinnate; pinnæ few, distant, narrow-linear, apiculate, the terminal one elongated, the lateral much shorter.—Hook, Ic. Plant. t. 960; Sp. Fil. iii. 35; Hook. f. Fl. Nov. Zel. ii. 31; Handb. N.Z. Fl. 369; Hook, and Bak. Syn. Fl. 181; Thoms. N.Z. Ferns, 69; Field, N.Z. Ferns, 110, t. 25, f. 4, 4A. Polybotrya nana, Fèe. Acrost. t. 38, f. 1, Blechnum nigrum, Mett.

North Island: Dark gloomy forests from Whangarei southwards, not common. South Island: Nelson—Collingwood, D. Grant; Takaka and West Wanganui, Kingsley. Westland — Abundant at low elevations, Enys! A. Hamilton! J. W. Brame! &c. Otago—Milford Sound, Bligh's Sound, Lyall, Hector and Buchanan. Sea-level to 3000 ft.

Easily recognised by the enlarged terminal portion of the frond, which is often only shallowly lobed, while the lower pinnæ are usually distinct from one another. The surface of the frond is often overgrown with mosses or hepaticæ, in the same manner as in Trichomanes elongatum.

12.L. fluviatilis, Spreng. Syst. Veg. iv. 65.—Rhizome stout, suberect, often woody, densely clothed with the bases of the old stipites and with chestnut-brown subulate scales. Stipes very short, densely scaly. Sterile fronds very numerous, forming a broad spreading crown at the top of the rhizome, l-2½ ft. high ¾-l ½ in. broad, linear or linear-lanceolate, submembranous, pale brownish-green, pinnate throughout; rhachis densely clothed with spreading subulate scales. Pinnæ very numerous, 20–50 pairs, ½–¾ in. long, ¼–⅓ in. broad, oblong to orbicular-oblong, obtuse, not decurrent, the lower more remote and often shortly stipitate, the upper sessile, the uppermost usually adnate; margins thin, sinuate or denticulate. Fertile fronds narrow-linear, erect; pinnæ ⅓–⅔ in. long, ⅛ in. broad, linear, obtuse, erecto-patent.—Hook. Sp. Fil. iii, 34; Hook.f. Fl. Nov. Zel. ii. 28; Fl. Tasm. ii. 142, t. 167; Handb. N.Z. Fl. 366; Hook, and Bak. Syn. Fil. 181; Benth. Fl. Austral. vii. 736; Thoms. N.Z. Ferns, 69; Field, N.Z. Ferns, 109, t. 27, f. 2, 2A. L. rotundifolia, Raoul, Choix, 9, t. 23; Col. in Tasmanian Journ. Nat. Sci. (1845) 19. Stegania fluviatilis, R. Br. Prodr. 152. Blechnum fluviatile, Mett.

North and South Islands, Chatham Islands, Stewart Island: From Hokianga and Whangaroa southwards, not uncommon in damp hilly forests. Sea-level to 2500 ft.

Also in Victoria and Tasmania. A crested form is occasionally seen, and has been described by Mr. Colenso as var. ramosa (Trans. N.Z. Inst. xx. 225).

page 984
13.L. membranacea, Col. ex Hook. Sp. Fl. iii. 34, t. 145.— Rhizome short, stout, suberect, clothed with the bases of the old stipites mixed with a few subulate scales. Stipes very short, scaly at the base. Fronds tufted at the top of the rhizome, the sterile ones 3–10 in. long, ¾–1 ½ in. broad, lanceolate or linear-lanceolate, acuminate, broadest above the middle, gradually narrowed to the base, rather membranous, pale - green, quite glabrous, pinnate; rhachis smooth, naked. Pinnæ numerous, spreading or erectopatent, the longest ½–¾ in. long, about ¼ in. broad, ovate-oblong or oblong, obtuse, broadly adnate at the base but not dilated nor decurrent, coarsely dentate-serrate, the lower quite distinct at the base, gradually becoming smaller and eventually reduced to mere rounded auricles, the uppermost more or less confluent. Fertile fronds usually longer than the sterile and with longer stipites, pinnate; Pinnæ distant, ⅙–½ in. long, linear, apiculate.—Hook. f. Handb. N.Z. Fl. 366; Hook, and Bak. Syn. Fil. 181; Thoms. N.Z. Ferns, 69; Field, N.Z. Ferns, 111, t. 5, f. 6, 6a. L. oligoneuron, Col. in Trans. N.Z. Inst. xvi. (1884) 346. L. intermedia, Col. l.c. xix. (1887) 274; L. pygmæa, Col. l.c. xxv. (1893) 322. Blechnum membranaceum. Mett.

North Island: Shaded places by the banks of streams, not uncommon throughout. South Island: In various localities along the east coast from Nelson to Otago, but apparently rare and local. Sea-level to 2000 ft.

Large forms of this, with longer and narrower pinnæ rather more closely placed, are difficult to distinguish from L. lanceolata, if, indeed, the two species do not pass directly into one another. In its usual state, however, it is a much smaller plant, with shorter and broader obtuse pinnæ, distinctly separated from one another, and not dilated at the base or decurrent as in lanceolata. I have seen no specimens from Canterbury or Otago.

14.L. Fraseri, A. Cunn. Precur. n. 185. — Rhizome erect, clothed with the bases of the old stipites and with a dense tuft of dark chestnut-brown scales at the tip, often elongated into a slender caudex 6–24 in. high or more, resembling the trunk of a miniature tree-fern. Stipes 3–9 in. high, scaly towards the base. Fronds forming a spreading crown at the top of the caudex, 9–18 in. long, 3–6 in. broad, ovate or ovate - oblong to ovate-lanceolate, acuminate, quite glabrous, almost membranous or subcoriaceous, bipinnate; rhachis with a narrow interrupted wing furnished with numerous triangular lobes. Pinnæ 2–3 in. long, ⅓–⅔ in. broad, lanceolate or linear-lanceolate, acuminate, cut down almost to the rhachis. Pinnules numerous, close-set, ¼–⅓ in. long, linear-oblong, somewhat falcate, acute or apiculate, entire or serrate. Veins indistinct, simple or forked. Fertile fronds similar to the sterile, but rather smaller and with narrower pinnules. Sori covering the whole under-surface.—Raoul, Choix, 37; Hook. Ic. Plant. t. 185; Sp. Fil. iii. 40; Hook. f. Fl. Nov. Zel. ii. 31; Handb. N.Z. Fl. 369; Hook, and Bak. Syn. Fil. 182; Thoms. N.Z. Ferns, 70; Field, N.Z. Ferns, 111, t. 24, f. 4, 4A. Blechnum Fraseri, Metten. page 985

North Island: Abundant in dry woods from the North Cape southwards to the Upper Waikato and Taranaki. South Island: Nelson—Massacre Bay, Lyall; West Wanganui, Kingsley; extending along the West Coast as far south as Charlestown, Kirk. Sea-level to 2000 ft.

A very handsome and distinct species, confined to New Zealand, unless a plant lately discovered in the Philippine Islands should prove to be the same.